The Cambridge natural history, Vol. 01 (of 10)Hickson, Sydney J. (Sydney John)
Science
The Cambridge natural history, Vol. 01 (of 10)
Hickson, Sydney J. (Sydney John)
Animals
Schaudinn has discovered and worked out true syngamic processes, some
bisexual, some exogamous, in several other Rhizopods. In _Chlamydophrys
stercorea_ the pairing-cells are equal, and are formed by the aggregation
of the chromidia into minute nuclei around which the greater part of the
cytoplasm aggregates, while the old nucleus (with a little cytoplasm) is
lost. These brood-cells are 2-flagellate pairing-cells, which are
exogamous: the zygote is a brown cyst; if this be swallowed by a mammal,
the original _Chlamydophrys_ appears in its faeces.[72]
_Centropyxis aculeata_, a species very common in mud or moss, allied to
_Difflugia_, also forms a brood by aggregation around nuclei derived from
chromidia. The brood-cells are amoeboid, and secrete hemispherical shells
like those of _Arcella_; some first divide into four smaller ones, before
secreting the shell. Pairing takes place between the large and the small
forms; and the zygote encysts. Weeks or months afterwards the cyst opens
and its contents creep out as a minute _Centropyxis_. Finally, _Amoeba
coli_ produces its zygote in a way recalling that of _Actinosphaerium_ (pp.
73-75, Fig. 21): the cell encysts; its nucleus divides, and each daughter
divides again into two, which fuse reciprocally. Thus the cyst contains two
zygote nuclei. After a time each of these divides twice, so that the mature
cyst contains eight nuclei. Probably when swallowed by another animal they
liberate a brood of eight young amoebae. Thus in different members of this
group we have exogamy, both equal and bisexual, and endogamy.
Most of the Rhizopoda live among filamentous Algae in pools, ponds, and in
shallow seas, etc.; some are "sapropelic" or mud-dwellers (many species of
_Amoeba_, _Pelomyxa_, _Difflugia_, etc.), others frequent the roots of
mosses. _Amoeba coli_ is often found as a harmless denizen of the large
intestine of man. _Amoeba histolytica_, lately distinguished therefrom by
Schaudinn, is the cause of tropical dysentery. It multiplies enormously in
the gut, and is found extending into the tissues, and making its way into
the abscesses that so frequently supervene in the liver and other organs.
_Chlamydophrys stercorea_ is found in the {58}faeces of several mammals.
The best monograph of this group is that of Penard.[73]
2. FORAMINIFERA[74]
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