The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
The abdomen bears no appendages. The telson is compressed in the
Calyptomera, and is produced into two flattened plates, one on each side
of the anal opening. The backwardly directed margins of these plates are
commonly serrated, and the lower corner of each is produced into a
curved spine, which carries secondary teeth. The number and arrangement
of these teeth, though often extremely variable in the same species, are
used extensively as specific characters. Above the anus the telson
commonly bears two long plumose hairs, which are directed backwards.
[Illustration:
FIG. 13.—_Bythotrephes cederströmii_, female, × 20, North Wales, from
a specimen found by A. D. Darbishire. _Car_, carapace.
]
In the Gymnomera the telson is not bilaterally compressed, and it may be
produced into a long spine, dorsal to the anus (e.g. _Bythotrephes_,
Fig. 13).
The =alimentary canal= is extremely simple. The labrum is large, and
forms a chamber above the mouth, into which food is driven by the limbs,
as in the Phyllopoda, food being taken while the animal swims or lies on
its back. The oesophagus runs vertically to join a small stomach, which
bends sharply backwards and passes gradually into an intestine. In the
last segment of the abdomen the intestine joins a short, thin-walled
rectum, provided with radial muscles, by means of which it can be
dilated. The dilatation of the rectum leads to an inhalation of water
through the anus, which may possibly serve as a means of respiration. In
the Daphniidae and Bosminidae there are two forwardly-directed digestive
glands which open into the stomach, and in _Eurycercus_ there is a large
caecum at the junction of the rectum with the intestine. The intestine
is usually straight, but in Lynceidae and in some Lyncodaphniidae it is
coiled (e.g. _Peracantha_, Fig. 14).
In _Leptodora_ the alimentary canal is altogether remarkable; the
oesophagus is a long and very narrow tube, which runs back through the
whole length of the thorax and joins the mid-gut in the third abdominal
segment. The mid-gut is not differentiated into stomach and intestine;
it has no diverticula of any kind, and runs straight backwards to join
the short rectum a little in front of the anus.
[Illustration:
FIG. 14.—_Peracantha truncata_, female, × 100. Oxford.
]
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