The Cambridge natural history, Vol. 05 (of 10)Sedgwick, Adam
Science
The Cambridge natural history, Vol. 05 (of 10)
Sedgwick, Adam
Animals
The _nervous system_ differs considerably from that in the Order
Chilognatha; it resembles that in the Schizotarsia, and differs again from
that in the other two Orders, Symphyla and Pauropoda. The brain shows some
differences from other Orders chiefly in the development of the different
lobes which are connected with the sense organs, the eyes and antennae, for
instance; but the most marked difference is in the ventral ganglionic cord.
First, the ganglionic swellings are much more clearly marked than in the
Chilognatha. Secondly, the first three ganglia differ {59}from the others
in being nearer to one another and forming a single mass when seen by the
naked eye, though when examined by the aid of a microscope we can see all
the different parts are there. Thirdly, the division into two cords
mentioned in the Chilognatha is carried to a much greater extent. The
ganglia in each segment can be seen plainly to be double, and the cords
connecting the ganglia are two in number. We can plainly see that the
ventral nervous system of the Chilopoda consists of two cords lying
parallel to one another, and each having a ganglionic enlargement in every
segment. Whether a visceral nervous system is present in the group is
doubtful.
The eighth family of the Chilognatha, the Polyxenidae, show an approach to
the Chilopod nervous system.
The generative system differs chiefly in the opening of the genital
apparatus at the end of the body instead of in the third segment; though
this difference only separates the Order from the Chilognatha and not from
the other Orders. They also have two pairs of large accessory glands (as
they are called) connected with the genital openings.
ORDER III. SCHIZOTARSIA.
The third Order of Myriapods, the Schizotarsia, show a much greater
resemblance to the Chilopoda than to the first Order, the Chilognatha.
There are, however, important differences to distinguish them from all the
other Orders.
The shape of the body is short, thick, and very compact. The composition of
the individual segments resembles that found in Chilopoda rather than that
of Chilognatha.
The antennae are very long, longer than in any of the Chilopods, and are
composed of a great number of very small joints. The mouth parts show a
greater length and slenderness than do those of the other Orders mentioned
as yet. They consist of—
1. An _upper lip_ partly free, but fused at the sides with the rest of
the head. The upper lip is in three parts, as in the Chilopoda, but with
the middle part very small and the lateral pieces large.
2. A pair of _jaws_ or _mandibles_. These are provided not only with
teeth, as in the other Myriapods, but also with a sort of comb of stiff
bristles.
{60}3 and 4. Two pairs of _maxillae_ or foot jaws distinguished by their
length and slenderness.
5. The _poison claws_ long, slender, and not sharply curved. The bases of
the poison claws hardly fused together and short.
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