The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
The kidneys consist of a large number of glomeruli, produced by the coiled
segmental tubes, each of which is composed of a nephrostome or funnel
opening into the body-cavity, a Malpighian body and an efferent canal. The
latter combine to form the segmental duct which opens into the cloaca. The
testes, composed of a large number of sperm-producing glands, are drained
by transverse canals which combine into a longitudinal canal, and this
again sends off numerous efferent canals which open into the efferent
canals of the kidney, so that the segmental duct (Leydig's duct of many
authors) conveys both sperma and urine.
{49}[Illustration: FIG. 7.–Diagrammatic representation of modifications of
the urino-genital ducts. 1, 2, Male and female Newt; 3, a tubule of the
kidney; 4, male _Rana_; 5, male _Bufo_; 6, male _Bombinator_; 7, male
_Discoglossus_; 8, male _Alytes_. _a_, Artery entering, and producing a
coil in, the Malpighian body, _M_; _B_, Bidder's organ; _ef.s.c_, efferent
segmental canal; _F.B_, fat-body; _gl_, glomerulus; _K_, kidney; _l.c.c_,
longitudinal collecting canal; _M_, Malpighian body; _Md_, Müllerian duct;
_N_, nephrostome; _O_, ovary; _Ov_, oviduct; _s.d_, segmental duct; _T_,
testis; _Ur_, ureter; _V.d_, vas deferens; _V.s_, vesicula seminalis.]
{50}In the female the network of transverse and longitudinal canals, which
originally connect the generative glands with the kidney's efferent canals,
is deduced in so far as the connection is interrupted and the vestiges of
the transverse canals are no longer functional. The eggs fall into the
body-cavity and are caught up by the ostium or inner abdominal opening of a
special duct, the oviduct (Müllerian duct of many authors). Vestiges, more
or less complete, of these oviducts persist in the males of most Amphibia.
This general scheme presents some modifications in the various groups of
Amphibia.
The Apoda retain the most primitive conditions. The kidneys are still long
and narrow, and the glomeruli are, at least in the anterior part of the
organ, still strictly segmental, agreeing in number and position, each with
a vertebral segment; later, the number of the glomeruli is greatly
increased, and the former agreement becomes quite disturbed. The generative
glands still retain their segmental arrangement, but they are restricted to
a much shorter region than the kidneys. In the male Apoda a considerable
portion of the cloaca can be everted by special muscles, and acts as an
intromittent organ. Both sexes possess a ventral urinary bladder.
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