But, if succession does not imply descent, descent, at least, implies
succession, and the fact that succession is the necessary corollary of
descent, may be used as a corrective for the erroneous allocations made
by neontologists on the basis of purely morphological considerations.
The _priority_ of a type is the _sine qua non_ condition of its
being accepted as _ancestral_. It is always embarrassing when, as
sometimes happens, a “descendant” turns out to be older than, or even
coëval with, his “ancestor.” If, however, the historical position of
a form can be made to coincide with its anatomical pretensions to
ancestry, then the inference of descent attains to a degree of logical
respectability that is impossible in the case of purely zoölogical
evidence. Recent years have witnessed a more drastic application of the
historical test to morphological speculations, and the result has been
a wholesale revision of former notions concerning phylogeny. “I could
easily,” says Bather, “occupy the rest of this hour by discussing the
profound changes wrought by this conception on our classification. It
is not that orders and classes hitherto unknown have been discovered,
not that some erroneous allocations have been corrected, but the whole
basis of our system is being shifted. So long as we were dealing with
a horizontal section across the tree of life—that is to say, with
an assemblage of approximately contemporaneous forms—or even with a
number of such horizontal sections, so long were we confined to simple
description. Any attempt to frame a causal connection was bound to be
speculative.” (_Ibidem_, p. 258.) Whether zoölogists will take kindly
to this “shifting of the whole basis” of classification, remains to be
seen. Personally, we think they would be very ill-advised to exchange
the solid observational basis of homology for the scanty facts and
fanciful interpretations of palæontologists.
The second stumbling block in the path of Transformism is the
occurrence of convergence. We have seen that, in the palæontological
argument, descent is inferred conjointly from similarity and
succession, and that, in the abstract, this argument is very
persuasive. One of the concrete phenomena, however, that tend to make
it inconsequential, is the undoubted occurrence of convergence. Prof.
H. Woods of Cambridge, in the Introduction to the 5th edition of his
“Palæontology” (1919), speaks of three kinds of convergence (cf.,
pp. 14, 15, 16), which, as a matter of convenience, we may term the
parallelistic, the radical, and the adaptational, types of convergence.
A brief description of each type will serve to elucidate its nature and
its significance:
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