But let us suppose the case for evolution to be ideally favorable, and
assume that in every instance we possessed a perfect gradation of forms
between two extremes, such, for example, as occurs in the Ammonite
series, even then we would be far from having a true demonstration of
the point at issue. Bateson has called our attention to the danger of
confounding sterile and instable _hybrids_ with intergradent species.
“Examine,” he says, “any two thoroughly distinct species which meet
each other in their distribution, as for instance, _Lychnis diurna_
and _vespertina_ do. In areas of overlap are many intermediate forms.
These used to be taken to be transitional steps, and the specific
distinctness of _vespertina_ and _diurna_ was on that account
questioned. Once it is known that these supposed intergrades are merely
mongrels between the two species the transition from one to the other
is practically beyond our powers of imagination to conceive. If both
these can survive, why has their common parent perished? Why, when
they cross, do they not reconstruct it instead of producing partially
sterile hybrids? I take this example to show how entirely the facts
were formerly misrepresented.” (_Heredity_, Smithson. Inst. Rpt. for
1915, p. 369.)
Similarly, T. H. Morgan has shown, with reference to _mutants_,
the fallacy of inferring common descent from the phenomenon of
intergradence, and what holds true for a series of intergradent mutants
would presumably also hold true of a series of intergradent species,
could such a series be found and critically distinguished from hybrid
and mutational intermediates. In short, the Darwinian deduction of
common origin from the existence of intergradence must now be regarded
as a thoroughly discredited argument. “Because we can often arrange
the series of structures in a line extending from the very simple
to the more complex, we are apt to become unduly impressed by this
fact and conclude that if we found the complete series we should find
all the intermediate steps and that they have arisen in the order
of their complexity. This conclusion is not necessarily correct.”
(“A Critique of the Theory of Evolution,” p. 9.) Having cited such
a series of gradational mutations ranging between the long-winged,
and completely wingless condition, in the case of the Vinegar Fly
(_Drosophila melanogaster_), as well as two similar graded series based
on pigmentation and eye color, he concludes: “These types, with the
fluctuations that occur within each type, furnish a complete series of
gradations; yet historically they have arisen independently of each
other. Many changes in eye color have appeared. As many as thirty or
more races differing in eye color are now maintained in our cultures.
Some of them are so similar that they can scarcely be separated from
each other. It is easily possible beginning with the darkest eye color,
sepia, which is a deep brown, to pick out a perfectly graded series
ending with pure white eyes.
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