The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and PlantsDawson, John William, Sir
Science
The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and Plants
Dawson, John William, Sir
Paleobotany; Paleontology
The supposed fern of the Upper Silurian, figured in the first edition of
this work, has proved on further examination to be merely an imitative
form produced by crystallisation. On the other hand, the recent
discovery of a cockroach and two species of Scorpion in the Silurian,
proves the existence of land animals as well as plants at this period.
[Illustration: FIG. 87.—_Psilophyton princeps_ (Dn.) Silurian and
Devonian. Restored.
_a_, Fruit, natural size. _b_, Stem, natural size. _c_, Scalariform
tissue of the axis, highly magnified. In the restoration one side is
represented in vernation, and the other in fruit.]
It is probable that these discoveries represent merely a small
proportion of the plants actually existing in the Silurian period. All
the deposits of this age at present known to us are marine; and most of
them were probably formed at a distance from land, so that it is little
likely that land plants could find their way into them. At any time the
discovery of an estuarine or lacustrine deposit of Silurian age might
wonderfully extend our knowledge of this ancient flora.
The Devonian or Erian age, that of the classic Old Red Sandstone of
Scotland, is that in which we find the first great and complete land
flora; and though this is inferior in number of species to that of the
succeeding Carboniferous, and greatly less important with reference to
its practical bearing on our welfare, it is in some respects superior in
that variety which depends on diversity of soil and of station. To
appreciate this, it will be necessary to glance at the range and
subdivisions of the modern flora.
In the modern world we divide all vegetation into two great series, that
of the Flowering Plants (_Phænogams_), which also produce true fruits
and seeds, and that of the Flowerless Plants (_Cryptogams_), which
produce minute spores instead of seeds. The latter is in every respect
the lower group. This lower series is again divisible into three
classes—first and lowest, that of the Seaweeds, Moulds, and Lichens
(Thallophytes). Secondly, that of the Mosses and their allies
(Anophytes). Thirdly, that of the Ferns, Equisetums and Club-mosses
(Acrogens). In like manner the second, or higher series is divisible
into three classes: that of the Pines and Cycads (Gymnosperms), having
naked seeds not covered by true fruits, and woody tissue of simple
structure; that of the Palms and Grasses and their allies (Endogens);
and last and highest, that of the ordinary timber trees and other plants
allied to them, with exogenous stems, netted-veined leaves, and a
two-leaved embryo (Exogens). These last are in every respect the
dominant plants on our present continents. Carrying with us this twofold
division of the vegetable kingdom and its subdivisions, we shall be
prepared to understand the relation of the more ancient floras to that
now living.
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