The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
On slightly pulling the two, so as to
imitate the action of the muscles, the three teeth will be found to
come together sharply, exactly in the manner described.
Works on mechanics are full of contrivances for the conversion of
motion; but it would, perhaps, be difficult to discover among these
a prettier solution of the problem; given a straight pull, how to
convert it into three simultaneous convergent movements of as many
points.
What I have called the _filter_ is constructed mainly out of
the chitinous lining of the pyloric chamber. The aperture of
communication between this and the cardiac chamber, already narrow,
on account of the constriction of the walls of the stomach at this
point, is bounded at the sides by two folds; while, from below, a
conical tongue-shaped process (figs. 6, 10, and 11, _cpv_), the
surface of which is covered with hairs, further obstructs the
opening. In the posterior half of the pyloric chamber, its side
walls are, as it were, pushed in; and, above, they so nearly meet in
the middle line, that a mere vertical chink is left between them;
while even this is crossed by hairs set upon the two surfaces. In
its lower half, however, each side wall curves outwards, and forms
a cushion-shaped surface (fig. 10, _cs_) which looks downwards and
inwards. If the {59} floor of the pyloric chamber were flat, a
wide triangular passage would thus be left open in its lower half.
But, in fact, the floor rises into a ridge in the middle, while, at
the sides, it adapts itself to the shape of the two cushion-shaped
surfaces; the result of which is that the whole cavity of the
posterior part of the pyloric division of the stomach is reduced to a
narrow three-rayed fissure. In transverse section, the vertical ray
of this fissure is straight, while the two lateral ones are concave
upwards (fig. 9, _E_). The cushions of the side walls are covered
with short close-set hairs. The corresponding surfaces of the floor
are raised into longitudinal parallel ridges, the edge of each of
which is fringed with very fine hairs. As everything which passes
from the cardiac sac to the intestine must traverse this singular
apparatus, only the most finely divided solid matters can escape
stoppage, so long as its walls are kept together.
Finally, at the opening of the pyloric sac into the intestine, the
chitinous investment terminates in five symmetrically arranged
processes, the disposition of which is such that they must play the
part of valves in preventing any sudden return of the contents of the
intestine to the stomach, while they readily allow of a passage the
other way. One of these valvular processes is placed in the middle
line above (figs. 10 and 11, _v_^1). It is longer than the others and
concave below. The lateral processes (_v_^2,) of which there are two
on each side, are triangular and flat. {60}
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