The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
The heart (fig. 15, _h_), lies in the midst of this sinus. It is a
thick muscular body (fig. 16), with an irregularly hexagonal contour
when viewed from above, one angle of the hexagon being anterior and
another posterior. The lateral angles of the hexagon are connected
by bands of fibrous tissue (_ac_) with the walls of the pericardial
sinus. Otherwise, the heart is free, except in so far as it is kept
in place by the arteries which leave it and traverse the walls of
the pericardium. One of these arteries (figs. 5, 12, and 16, _saa_),
starting from the hinder part of the heart, of which it is a sort of
continuation, runs along the middle line of the abdomen above the
intestine, to which it gives off many branches. A second large artery
starts from a dilatation, which is common to it with the foregoing,
but passing directly downwards (figs. 12 and 15, _sa_, and fig. 16,
_st.a_), either on the right or on the left side of the intestine,
{72} traverses the nervous cord (figs. 12 and 15), and divides into
an anterior (fig. 12, _sa_) and a posterior (_iaa_) branch, both of
which run beneath and parallel with that cord. A third artery runs,
from the front part of the heart, forwards in the middle line, over
the stomach, to the eyes and fore part of the head (figs. 5, 12, and
16, _oa_); and two others diverge one on each side of this, and sweep
{73} round the stomach to the antennæ (_aa_). Behind these, yet two
other arteries are given off from the under side of the heart, and
supply the liver (_ha_). All these arteries branch out and eventually
terminate in fine, so-called _capillary_, ramifications.
[Illustration: FIG. 16.—_Astacus fluviatilis._—The heart (× 4). A,
from above; B, from below; C, from the left side. _aa_, antennary
artery; _ac_, alæ cordis, or fibrous bands connecting the heart with
the walls of the pericardial sinus; _b_, bulbous dilatation at the
origin of the sternal artery; _ha_, hepatic artery; _la_, lateral
valvular apertures; _oa_, ophthalmic artery; _s.a_, superior valvular
apertures; _s.a.a_, superior abdominal artery; _st.a_, sternal
artery, in B cut off close to its origin.]
In the dorsal wall of the heart two small oval apertures are visible,
provided with valvular lips (fig. 16, _sa_), which open inwards, or
towards the internal cavity of the heart. There is a similar aperture
in each of the lateral faces of the heart (_la_), and two others in
its inferior face (_ia_), making six in all. These apertures readily
admit fluid into the heart, but oppose its exit. On the other hand,
at the origins of the arteries, there are small valvular folds,
directed in such a manner as to permit the exit of fluid from the
heart, while they prevent its entrance.
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