As to the structure of the vitreous body, apart from the fibres and
pigment streaks already mentioned, I find it to be made up of prisms
extending from retina to capsule of lens, each containing a central
axis or fibre. Fig. 64 shows that the space around the pigment areas
and “visual fibres,” instead of being homogeneous, is wholly filled
with the polygonal cross-sections of these prisms. In Charybdea they
are generally more difficult to perceive than in my best material of
Tripedalia which was killed in acetic acid. In this the polygonal areas
stood apart from each other more plainly. Curiously enough I have been
unable to demonstrate in Tripedalia the “visual fibres” of Schewiakoff.
Here and there were found spaces that at first sight reminded of them
(Fig. 69, _sh_), but they contained no central fibre, and were probably
due to shrinkage. The polygonal areas themselves, however, often
contained a clear spot in the centre, at one side of which would be found
the cross-section of the fibre, as is shown in many cases in Fig. 68.
The clear spot is here undoubtedly due to shrinkage of the gelatinous
substance of the prism.
I think that these prisms and fibres are the direct continuations of
retinal cells. In a section such as that drawn in Fig. 63, which takes
just the very tops of the cells of a slightly pigmented retina, in the
centre of the section just grazing the space that lies between the retina
and the shrunken vitreous body, most of the cells toward the middle
(where especially the extreme tips are taken) show in their centres a
dot exactly corresponding to the dots in the polygonal areas of the
vitreous body. In the exact middle of the section, where only the cell
walls appear, slightly indicated, a dot is seen in each case. The size
and shape of the ends of the cells correspond with those of the polygonal
areas in the vitreous body, and I do not doubt that the latter are
continuations of the former. The vitreous body, then, instead of being
homogeneous, is composed of the clear highly refracting outer ends of
retinal cells. The assumption lies near that these are the true visual
rods, but of course it is assumption only.
To give a brief review, the points in which my conclusions differ from
those of Schewiakoff are as follows: I find (1) that the long pigment
streaks are parts of retinal cells continued into processes like his
visual rods; (2) that the vitreous body is composed of prisms with
central fibres proceeding from retinal cells; (3) that I am unable to get
satisfactory evidence of two types of cell distinguishable within the
retina, and at any rate find considerable evidence against the two types
he distinguishes.
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