The Descent of Man and Selection in Relation to Sex, Vol. II (1st Edition)Darwin, Charles
Science
The Descent of Man and Selection in Relation to Sex, Vol. II (1st Edition)
Darwin, Charles
Evolution (Biology); Human beings -- Origin; Sex differences; Sexual dimorphism (Animals); Sexual selection in animals
The case of the Argus pheasant is still more striking. The immensely
developed secondary wing-feathers, which are confined to the male, are
ornamented with a row of from twenty to twenty-three ocelli, each above
an inch in diameter. The feathers are also elegantly marked with oblique
dark stripes and rows of spots, like those on the skin of a tiger and
leopard combined. The ocelli are so beautifully shaded that, as the Duke
of Argyll remarks,[143] they stand out like a ball lying loosely within
a socket. But when I looked at the specimen in the British Museum, which
is mounted with the wings expanded and trailing downwards, I was greatly
disappointed, for the ocelli appeared flat or even concave. Mr. Gould,
however, soon made the case clear to me, for he had made a drawing of a
male whilst he was displaying himself. At such times the long secondary
feathers in both wings are vertically erected and expanded; and these,
together with the enormously elongated tail-feathers, make a grand
semicircular upright fan. Now as soon as the wing-feathers are held in
this position, and the light shines on them from above, the full effect
of the shading comes out, and each ocellus at once resembles the
ornament called a ball and socket. These feathers have been shewn to
several artists, and all have expressed their admiration at the perfect
shading.
It may well be asked, could such artistically-shaded ornaments have been
formed by means of sexual selection? But it will be convenient to defer
giving an answer to this question until we treat in the next chapter of
the principle of gradation.
The primary wing-feathers, which in most gallinaceous birds are
uniformly coloured, are in the Argus pheasant not less wonderful objects
than the secondary wing-feathers. They are of a soft brown tint with
numerous dark spots, each of which consists of two or three black dots
with a surrounding dark zone. But the chief ornament is a space parallel
to the dark-blue shaft, which in outline forms a perfect second feather
lying within the true feather. This inner part is coloured of a lighter
chesnut, and is thickly dotted with minute white points. I have shewn
this feather to several persons, and many have admired it even more than
the ball-and-socket feathers, and have declared that it was more like a
work of art than of nature. Now these feathers are quite hidden on all
ordinary occasions, but are fully displayed when the long secondary
feathers are erected, though in a widely different manner; for they are
expanded in front like two little fans or shields, one on each side of
the breast near the ground.
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