The Elements of Botany, For Beginners and For SchoolsGray, Asa
Science
The Elements of Botany, For Beginners and For Schools
Gray, Asa
Plants
254. The flowers of Larkspur, and of Monkshood or Aconite, which are
nearly related, are both strikingly irregular in calyx and corolla, and
considerably unsymmetrical. In Larkspur (Fig. 239-241) the irregular
calyx consists of five sepals, one of which, larger than the rest, is
prolonged behind into a large sac or spur; but the corolla is of only
four petals (of two shapes),--the fifth, needed to complete the
symmetry, being left out. And the Monkshood (Fig. 242-244) has five very
dissimilar sepals, and a corolla of only two very small and
curiously-shaped petals,--the three needed to make up the symmetry being
left out. The stamens in both are out of symmetry with the ground-plan,
being numerous. So are the pistils, which are usually diminished to
three, sometimes to two or to one.
255. =Flowers with Multiplication of Parts= are very common. The stamens
are indefinitely numerous in Larkspur and in Monkshood (Fig. 242, 243),
while the pistils are fewer than the ground-plan suggests. Most
Cactus-flowers have all the organs much increased in number (Fig. 229),
and so of the Water-Lily. In Anemone (Fig. 233) the stamens and pistils
are multiplied while the petals are left out. In Buttercups or Crowfoot,
while the sepals and petals conform to the ground-plan of five, both
stamens and pistils are indefinitely multiplied (Fig. 245).
[Illustration: Fig. 245. Flower of Ranunculus bulbosus, or Buttercup, in
section.]
256. =Flowers modified by Union of Parts=, so that these parts more or
less lose the appearance of separate leaves or other organs growing out
of the end of the stem or receptacle, are extremely common. There are
two kinds of such union, namely:--
_Coalescence_ of parts of the same circle by their contiguous margins;
and
_Adnation_, or the union of adjacent circles or unlike parts.
257. =Coalescence= is not rare in leaves, as in the upper pairs of
Honeysuckles, Fig. 163. It may all the more be expected in the crowded
circles or whorls of flower-leaves. Datura or Stramonium (Fig. 246)
shows this coalescence both in calyx and corolla, the five sepals and
the five petals being thus united to near their tips, each into a tube
or long and narrow cup. These unions make needful the following terms:--
_Gamopetalous_, said of a corolla the petals of which are thus
coalescent into one body, whether only at base or higher. The union may
extend to the very summit, as in Morning Glory and the like (Fig. 247),
so that the number of petals in it may not be apparent. The old name for
this was _Monopetalous_, but that means "one-petalled;" while
gamopetalous means "petals united," and therefore is the proper term.
[Illustration: Fig. 246. Flower of Datura Stramonium; gamosepalous and
gamopetalous.]
[Illustration: Fig. 247. Funnelform corolla of a common Morning Glory,
detached from its polysepalous calyx.]
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