The Elements of Botany, For Beginners and For SchoolsGray, Asa
Science
The Elements of Botany, For Beginners and For Schools
Gray, Asa
Plants
[Illustration: Fig. 299. Stamen of Globe Amaranth; very short filament
bearing a single anther-cell; it is open from top to bottom, showing the
pollen within.]
[Illustration: Fig. 300-305. Stamens of several plants of the Labiate or
Mint Family. Fig. 300. Of a Monarda: the two anther-cells with bases
divergent so that they are transverse to the filament, and their
contiguous tips confluent, so as to form one cell opening by a
continuous line. Fig. 301. Of a Calamintha: the broad connective
separating the two cells. Fig. 302. Of a Sage (Salvia Texana); with long
and slender connective resembling forks of the filament, one bearing a
good anther-cell; the other an abortive or poor one. Fig. 303. Another
Sage (S. coccinea), with connective longer and more thread-shaped, the
lower fork having its anther-cell wholly wanting. Fig. 304. Of a White
Sage, Audibertia grandiflora; the lower fork of connective a mere
vestige. Fig. 305. Of another White Sage (A. stachyoides), the lower
fork of connective suppressed.]
291. But anthers may become _one-celled_, and that either by confluence
or by suppression.
292. By confluence, when the two cells run together into one, as they
nearly do in most species of Pentstemon (Fig. 297), more so in Monarda
(Fig. 300), and completely in the Mallow (Fig. 298) and all the Mallow
family.
293. By suppression in certain cases the anther may be reduced to one
cell or halved. In Globe Amaranth (Fig. 299) there is a single cell
without vestige of any other. Different species of Sage and of the White
Sages of California show various grades of abortion of one of the
anther-cells, along with a singular lengthening of the connective (Fig.
302-305).
294. The splitting open of an anther for the discharge of its pollen is
termed its _Dehiscence_.
[Illustration: Fig. 306. Stamen with the usual dehiscence of anther down
the side of each cell.]
[Illustration: Fig. 307. Stamen of Pyrola; cells opening by a terminal
hole.]
[Illustration: Fig. 308. Stamen of Barberry; cells of anther each
opening by an uplifted valve.]
295. As the figures show, this is commonly by a line along the whole
length of each cell, either lateral or, when the anthers are extrorse,
often along the outer face, and when introrse, along the inner face of
each cell. Sometimes the opening is only by a chink, hole, or pore at
the top, as in the Azalea, Pyrola (Fig. 307), etc.; sometimes a part of
the face separates as a sort of trap-door (or valve), hinged at the top,
and opening to allow the escape of the pollen, as in the Sassafras,
Spice-bush, and Barberry (Fig. 308).
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