protectively coloured under side of the wing is not turned towards the
light but away from it. Moreover, in the pupa the anterior wings cover
the posterior ones completely, no matter what the wing position may be
later in the perfect insect. Furthermore, the thick and often darkly
coloured sheath of the pupa prevents the light having any effect, and
not a few species pass their pupal stage in such dark places--for
instance, under stones, as in the case of many 'Blues'--that the light
can hardly reach them. And if the light did exercise an influence, how
could it produce such diverse coloration as the protective colours of
diurnal butterflies, on the one side dark, even to blackness, on the
other side, yellow, reddish, and even white and pure green; and how
should the same rays of light call forth complicated colour patterns
on one and the same surface, for instance, the white, sprinkled with
green, of the Aurora butterfly (_Anthocharis cardaminis_)? Finally, we
have only to remember that numerous nocturnal Lepidoptera pass through
their pupa stage underground, although they exhibit brilliant as well
as protective colours in the most appropriate distribution, to reject
once for all the hypothesis that the influence of light plays any
decisive rôle in determining the distribution of the colours on the
wings of Lepidoptera.
But it is otherwise with _Tropidoderus_. In this case the wings grow
gradually during the slow growth of the animal, which takes place in
full light, and the wings of the young insect probably lie one above
the other, in exactly the same position, and cover the same places as
in the full-grown form; we might, therefore, from the facts of the
case, admit the possibility that the yellow of the covered portions is
due to the exclusion of light.
[Illustration: FIG. 11. _Tropidoderus childreni_, after Brunner von
Wattenwyl, in flying pose. _V_ anterior wing. _H. häut_, membranous
part of posterior wing. _H. horn_, horny portion.]
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