If we think of the proposition we arrived at earlier, and which was
proved chiefly by the case of identical twins, that the facies or
'type' of the descendant is determined at fertilization, we may be
inclined to regard such an oscillation of the hereditary tendencies
as almost impossible, for it means that, with the given mingling of
parental germ-plasms, the potency of inheritance from the two parents
in every part of the offspring is determined once for all in advance.
But the case of identical twins corroborates these oscillations, for
in them, too, the father predominates in one part, the mother in
another, and it proves, at the same time, that these oscillations do
not depend on any chances whatever in development, but that they are
exactly predetermined in the mingling of the hereditary substances in
the germ-plasm of the fertilized ovum, and are strictly adhered to
throughout development.
This fact can only be explained thus: the primary constituents of
the different parts and characters of the body are contained in the
parental germ-plasm in varying degrees of hereditary or transmissive
strength, and this can be understood very well from our point of view
without putting anything new into the 'portmanteau' of our theory
(Delage).
But I must digress a little in order to make this plain.
When, in speaking of plant-hybrids, I said that the collective ids
of the germ-plasm of a species must be equivalent in regard to the
characters of the species, I did not speak quite precisely; in the
majority of ids, in many cases in an overwhelming majority, this must
be the case, but not actually in all, at least not on the assumption
we make that the transformation of species is accomplished under the
control of natural selection.
Public-domain text, read in full here on John Shaqi.
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