zoological gardens to undertake such experiments from the point of view
of the germ-plasm theory, and their success would afford material for
the criticism of the theory, the more valuable because it is apparent
from the experiments on plants that the processes of heredity are
manifold, and are far from being uniform in different domains[10].
[10] Castle and Allen have recently published the results of
experiments in crossing white mice with grey, and these confirm
Mendel's Law.
I have assumed for my theory that the reducing division took place
according to the laws of chance, and that thus every combination of ids
occurred with equal frequency. This assumption seems to be confirmed,
by the experiments of the botanists I have mentioned, only in so far
that in the crossing of hybrids with one another every combination
of distinctive characters occurred with equal frequency. But, on the
other hand, the splitting of the germ-plasm at the reducing division
seems, as I said before, in many cases to take place in such a way that
the id-groups of the two parents are discretely separated from one
another; this was so in the stocks, peas, beans, and other hybrids.
But even if this were always the case in these, we could hardly infer
that it must be the same everywhere; we should rather expect that the
relationship of the two parents and their ids would bring into play the
finer attractions and repulsions between the ids of the germ-plasm,
and would thus determine their arrangement and grouping. Further
investigations may clear up this point; in the meantime we can only
say that already--even among hybrids--many deviations from Mendel's Law
have been established, for instance, by Bateson and Saunders (1902).
Before I conclude this lecture I should like to refer briefly to a
phenomenon which Darwin was acquainted with and sought to explain
through his theory of Pangenesis, but which at a later date was
regarded as not sufficiently authenticated to justify any attempt
at a theoretical explanation, since it seemed to contradict all our
conceptions of hereditary substance and its operations. I refer to the
phenomenon to which the botanists have given the pretty name of Xenia
(guest-gifts), and which consists in the fact that in crosses of two
different plants the characters of the male may appear not only in the
young plant but even in the seed, so that a transference of paternal
characters seems to take place from the pollen-tube to the mother, to
the 'tissue of the maternal ovary.' In heads of yellow-grained maize
(_Zea_) it is said that, after dusting with pollen from a blue-seeded
variety, blue seeds appear among the yellow, and similar observations
on other cultivated plants have been on record for more than half a
century. Thus dusting the stigma of green varieties of grape with the
pollen of a dark blue kind is said frequently to give rise to dark blue
fruits.
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