The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surfaceWallace, Alfred Russel
Science
The Geographical Distribution of Animals, Volume 1: With a study of the relations of living and extinct faunas as elucidating the past changes of the Earth's surface
Wallace, Alfred Russel
Paleontology; Zoogeography
The divisions in use till quite recently were of two kinds; either those
ready made by geographers, more especially the quarters or continents of
the globe; or those determined by climate and marked out by certain
parallels of latitude or by isothermal lines. Either of these methods was
better than none at all; but from the various considerations explained in
the preceding chapters, it will be evident, that such divisions must have
often been very unnatural, and have disguised many {53}of the most
important and interesting phenomena which a study of the distribution of
animals presents to us.
The merit of initiating a more natural system, that of determining
zoological regions, not by any arbitrary or _a priori_ consideration but by
studying the actual ranges of the more important groups of animals, is due
to Mr. Sclater, who, in 1857, established six primary zoological regions
from a detailed examination of the distribution of the chief genera and
families of Birds. Before stating what these regions are, what objections
have been made to them, what other divisions have been since proposed, and
what are those which we shall adopt in this work, it will be well to
consider the general principles which should guide us in the choice between
rival systems.
_Principles on which Zoological Regions should be formed._--It will be
evident in the first place that nothing like a perfect zoological division
of the earth is possible. The causes that have led to the present
distribution of animal life are so varied, their action and reaction have
been so complex, that anomalies and irregularities are sure to exist which
will mar the symmetry of any rigid system. On two main points every system
yet proposed, or that probably can be proposed, is open to objection; they
are,--1stly, that the several regions are not of equal rank;--2ndly, that
they are not equally applicable to all classes of animals. As to the first
objection, it will be found impossible to form any three or more regions,
each of which differs from the rest in an equal degree or in the same
manner. One will surpass all others in the possession of peculiar families;
another will have many characteristic genera; while a third will be mainly
distinguished by negative characters. There will also be found many
intermediate districts, which possess some of the characteristics of two
well-marked regions, with a few special features of their own, or perhaps
with none; and it will be a difficult question to decide in all cases which
region should possess this doubtful territory, or whether it should be
formed into a primary region itself. Again, two regions which have now
well-marked points of difference, may be shown to have been much more alike
at a comparatively recent geological epoch; {54}and this, it may be said,
proves their fundamental unity and that they ought to form but one primary
region. To obviate some of these difficulties a binary or dichotomous
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