The Geological History of PlantsDawson, John William, Sir
Science
The Geological History of Plants
Dawson, John William, Sir
Paleobotany
The most ancient of these contrivances, and one still continued in the
world of plants, is that of the barred or scalariform vessel. This may
be either square or hexagonal, so as to admit of being packed without
leaving vacancies. It is strengthened by a thick bar of ligneous matter
up each angle, and these are connected by cross-bars so as to form a
framework resembling several ladders fastened together. Hence the name
_scalariform_, or ladder-like. Now, in a modern Lycopod there is a
central axis of such barred vessels associated with simpler fibres or
elongated cells. Even in _Sphenophyllum_ and _Psilophyton_, already
referred to as allied to Rhizocarps,[AV] there is such a central axis,
and in the former rigidity is given to this by the vascular and woody
elements being arranged in the form of a three-sided prism or three-rayed
star. But such arrangements would not suffice for a tree, and hence
in the arboreal Lycopods of the Erian age a more complex structure is
introduced. The barred vessels were expanded in the first instance into
a hollow cylinder filled in with pith or cellular tissue, and the outer
rind was strengthened with greatly thickened cells. But even this was not
sufficient, and in the older stems wedge-shaped bundles of barred tissue
were run out from the interior, forming an external woody cylinder, and
inside of the rind were placed bundles of tough bast fibres. Thus, a stem
was constructed having pith, wood, and bark, and capable of additions to
the exterior of the woody wedges by a true exogenous growth. The plan
is, in short, the same with that of the stems of the exogenous trees of
modern times, except that the tissues employed are less complicated.
The structures of these remarkable trees, and the manner in which
they anticipate those of the true exogens of modern times, have been
admirably illustrated by Dr. Williamson, of Manchester. His papers, it
is true, refer to these plants as existing in the Carboniferous age, but
there is every reason to believe that they were of the same character
in the Erian. The plan is the same with that now seen in the stems of
exogenous phænogams, and which has long ceased to be used in those of the
Lycopods. In this way, however, large and graceful lycopodiaceous trees
were constructed in the Erian period, and constituted the staple of its
forests.
[AV] First noticed by the author, "Journal of Geological Society," 1865;
but more completely by Renault, "Comptes Rendus," 1870.
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