The Life and Letters of George John Romanes, M.A., LL.D., F.R.S.Romanes, George John
Science
The Life and Letters of George John Romanes, M.A., LL.D., F.R.S.
Romanes, George John
Naturalists -- Great Britain -- Biography; Romanes, George John, 1848-1894
[7] The following extract from 'An Examination of Weismannism,' pp.
2, 3, will possibly explain the theory of Pangenesis, which assumes:
1. That all the component cells of a multicellular organism throw off
inconceivably minute germs, or 'gemmules,' which are then dispersed
throughout the whole system.
2. That these gemmules, when so dispersed and supplied with proper
nutriment, multiply by self-division, and, under suitable conditions,
are capable of developing into physiological cells like those from
which they were originally and severally derived.
3. That, while still in this gemmular condition, these cell-seeds
have for one another a mutual affinity, which leads to their being
collected from all parts of the system by the reproductive glands of
the organism; and that, when so collected, they go to constitute the
essential material of the sexual elements--ova and spermatozoa being
thus aggregated packets of gemmules, which have emanated from all the
cells of all the tissues of the organism.
4. That the development of a new organism out of the fusion of
two such packets of gemmules is due to a summation of all the
developments of some of the gemmules which these two packets contain.
5. That a large proportional number of the gemmules in each packet,
however, fail to develop, and are then transmitted in a dormant
state to future generations, in any of which they may be developed
subsequently, thus giving rise to the phenomena of reversion or
atavism.
6. That in all cases the development of gemmules into the form
of their parent cells depends on their suitable union with other
partially developed gemmules which precede them in the regular course
of growth.
7. That gemmules are thrown off by all physiological cells, not only
during the adult state of the organism, but during all stages of
its development. Or, in other words, that the production of these
cell-seeds depends upon the adult condition of parent cells, not upon
that of the multicellular organism as a whole.
[8] The experiments in graft-hybridisation were to prove that
formative material (or gemmules) was actually present in the general
tissues of plants and was capable of uniting with the gemmules of
another plant and thus of reproducing the entire organism. For if
the hybrid, afterwards produced, presents equally the characters
of the scion and the stock, then formative material must have been
present in the tissues of the scion, and it is demonstrated that the
somatic tissues of the scion have exercised an effect on the germinal
elements of the stock, inasmuch as it has caused their offspring in
part to resemble it. Such facts Romanes considered to be fully in
harmony with the theory of Pangenesis, and inconsistent with any
theory which supposes that _no_ part of the parent organism generates
_any_ of the formative material.
[9] To the Royal Society.
[10] For the Suppression of Vivisection.
[11] Mr. Grant Allen.
[12] Spiritualism.
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