I have had it twice at
different times and from different growers.” Similarly, the long-headed
green dahlia arose twice over some years ago in the nursery of Messrs
Zocher & Co.
Further, the peloric Toad-flax (_Linaria vulgaris peloria_) is, De Vries
informs us, “known to have originated from the ordinary type at different
times and in different countries under more or less divergent
conditions.” And, as this variety is wholly barren, it must in each
instance have had an independent origin. Lastly, the purple beech seems
to be a mutation which has originated at least three times over.
Mutation Theory Criticised
Every one interested in biological theory should read both _Species and
Varieties_ and _Plant Breeding_ by De Vries, works which are of
incalculable value to the horticulturist and agriculturist as well as to
the biologist.
While not wishing to detract in any way from the truly splendid work done
by De Vries, we feel constrained to bring several charges against him.
Firstly, he suffers from the complaint that seizes nine out of ten
originators of new theories. He pushes his theory to extreme lengths; he
allows his imagination to run away with him. We do not think that on the
evidence available he is justified in asserting that every species passes
through alternating periods of comparative quiescence and periods in
which it throws off, as mutations, swarms of elementary species. He is
justified in asserting that discontinuous variation is by no means an
uncommon phenomenon, but further than this it does not seem safe to go at
present.
Secondly, he ought to lay more stress on the fact that _Oenothera
lamarckiana_ is a plant which does not appear to be known in the wild
state, and that it is therefore possibly a hybrid plant, and the
so-called elementary species which it gives off may be merely the
varieties out of which it has been built up. Boulenger and Bailey have
both studied this plant, and they have not been able to witness all the
mutations of which De Vries speaks, so that the former says, “The fact
that _Oenothera lamarckiana_ was originally described from a garden
flower, grown in the Paris _Jardin des Plantes_, and that, in spite of
diligent search, it has not been discovered wild anywhere in America,
favours the probability that it was produced by crossing various forms of
the polymorphic _Oenothera biennis_, which had been previously introduced
in Europe.”
Definition of a Species
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