Let us now combine the processes of plant and animal; we start with
the latter. In it we have a mechanism which does work. The source of
its energy is the potential chemical energy of its foodstuffs, which
latter reduce down to those substances known as proteids, fats, and
carbohydrates. The energy-value of these compounds is considerable,
that is to say, if they are burned in a stream of oxygen a large
quantity of heat is obtained from their combustion. They are ingested
by the animal, broken down chemically, and rearranged. The proteids
eaten by the animal (say those of beef or mutton or wheat) are acted
upon by the enzymes of the alimentary canal and are decomposed into
their immediate constituents, amino-acids, and then other enzymes
rearrange these amino-acids so as to form proteid again, but proteids
of the same kinds as those characteristic of the tissues. This
decomposition and re-synthesis is carried out also with respect to the
fats and carbohydrates ingested. The result is that the food taken into
the alimentary canal, or at least a part of it, is built up into the
living substance of the animal’s body. The energy expended upon these
processes of digestion and assimilation is probably inconsiderable.
During these processes the animal absorbs available chemical energy.
The energy thus taken into the animal is then transformed. The major
part of it appears as mechanical energy--that of bodily movement, the
movements of heart, lungs, blood, etc.--and heat. Some part of it
becomes nervous energy, by which rather vague term we mean the energy
involved in the propagation of nervous impulses. Some of it is used
in glandular reactions, in the formation of the digestive juices, for
instance. The most of it, however, transforms to mechanical energy
and heat. Just how these energy transformations are effected we do
not know. The heat is, of course, the result of chemical changes,
oxidations, decompositions, or changes of the same kind as that of the
dilution of sulphuric acid by water, but the mechanical energy appears
to result directly from chemical change without the intermediation of
heat. We shall return to this point in a later chapter, and content
ourselves with saying here that the chemical compounds contained in
the metabolic tissues of the animal body undergo transformation from
a state of high to a state of low chemical potential, and that this
difference of potential is represented by the work done and the heat
generated. The proteid, fat, and carbohydrate of the tissues represent
the condition of high potential; and the carbon dioxide, the water, and
the urea, into which these substances are transformed, represent the
condition of low potential.
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