Botany; Geotropism; Plants -- Irritability and movements
When a radicle is placed in a horizontal or inclined position, the
terminal growing part, as is well known, bends down towards the centre
of the earth; and Sachs[5] has shown that whilst thus bending, the
growth of the lower surface is greatly retarded, whilst that
of the upper surface continues at the normal rate, or may be even
somewhat increased. He has further shown by attaching a thread, running
over a pulley, to a horizontal radicle of large size, namely that of
the common bean, that it was able to pull up a weight of only one
gramme, or 15.4 grains. We may therefore conclude that geotropism does
not give a radicle force sufficient to penetrate the ground, but merely
tells it (if such an expression may be used) which course to pursue.
Before we knew of Sachs’ more precise observations we covered a flat
surface of damp sand with the thinnest tin-foil which we could procure
(.02 to .03 mm., or .00012 to .00079 of an inch in thickness), and
placed a radicle close above, in such a position that it grew almost
perpendicularly downwards. When the apex came into contact with the
polished level surface it turned at right angles and glided over it
without leaving any impression; yet the tin-foil was so flexible, that
a little stick of soft wood, pointed to the same degree as the end of
the radicle and gently loaded with a weight of only a quarter of an
ounce (120 grains) plainly indented the tin-foil.
[5] ‘Arbeiten des bot. Inst. Würzburg,’ vol. i. 1873, p. 461. See also
p. 397 for the length of the growing part, and p. 451 on the force of
geotropism.
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