The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
for the sap, and elsewhere into other components of the general structure.
The many changes implied are effected in various ways: now by that simple
re-arrangement of components known as isomeric change; now by that taking
from a compound one of its elements and inserting one of another kind,
which is known as substitution; and now by oxidation, as when the
oxy-cellulose which constitutes wood-fibre, is produced.
Besides elaborating building materials, the protoplasm elaborates
itself--that is, elaborates more of itself. It is chemically distinguished
from the building materials by the presence of nitrogen. Derived from
atmospheric ammonia, or from decaying or excreted organic matter, or from
the products of certain fungi and microbes at its roots, the nitrogen in
one or other combination is brought into a plant by the upward current; and
by some unknown process (not dependent on light, since it goes on equally
well if not better in darkness) the protoplasm dissociates and appropriates
this combined nitrogen and unites it with a carbo-hydrate to form one or
other proteid--albumen, gluten, or some isomer; appropriating at the same
time from certain of the earth-salts the requisite amount of sulphur and in
some cases phosphorus. The ultimate step, as we must suppose, is the
formation of living protoplasm out of these non-living proteids. A cardinal
fact is that proteids admit of multitudinous transformations; and it seems
not improbable that in protoplasm various isomeric proteids are mingled. If
so, we must conclude that protoplasm admits of almost infinite variations
in nature. Of course _pari passu_ with this dual process--augmentation of
protoplasm and accompanying production of carbo-hydrates--there goes
extension of plant-structure and plant-life.
To these essential metabolic processes have to be added certain ancillary
and non-essential ones, ending in the formation of colouring matters,
odours, essential oils, acrid secretions, bitter compounds and poisons:
some serving to attract animals and others to repel them. Sundry of these
appear to be excretions--useless matters cast out, and are doubtless
katabolic.
The relation of these facts here sketched in rude outline to the doctrine
of Evolution at large should be observed. Already we have seen how (§ 8a),
in the course of terrestrial evolution, there has been an increasingly
heterogeneous assemblage of increasing heterogeneous compounds, preparing
the way for organic life. And here we may see that during the development
of plant-life from its lowest algoid and fungoid forms up to those forms
which constitute the chief vegetal world, there has been an increasing
number of complex organic compounds formed; displayed at once in the
diversity of them contained in the same plant and in the still greater
diversity displayed in the vast aggregate of species, genera, orders, and
classes of plants.
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