The Principles of Biology, Volume 2 (of 2) — John Shaqi
The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
it shows the older sheaths in process of being split open. That is to
say, there must result the mode of growth which helped to give the name
Endogens to this class.
[Illustration: Figs. 92–94.]
[Illustration: Figs. 95–99.]
The other way in which an integrated series of fronds may acquire
the rigidity needful for maintaining an erect position, has next to
be considered. If the successive fronds do not acquire such habit
of curling as may be taken advantage of by natural selection, so as
to produce the requisite stiffness; then, the only way in which the
requisite stiffness appears producible, is by the thickening and
hardening of the fused series of mid-ribs. The incipient axis will not,
in this case, be inclosed by the rolled-up fronds; but will continue
exposed. Survival of the fittest will favour the genesis of a type, in
which those portions of the successive mid-ribs that enter into the
continuous bond, become more bulky than the disengaged portions of the
mid-ribs: the individuals which thrive and have the best chances of
leaving offspring, being, by the hypothesis, individuals having axes
stiff enough to raise their foliage above that of their fellows. At
the same time, under the same influences, there will tend to result an
elongation of those portions of the mid-ribs, which become parts of the
incipient axis; seeing that it will profit the plant to have its leaves
so far removed from one another, as to prevent mutual interferences.
Hence, from the recumbent type there will evolve, by indirect
equilibration (§ 167), such modifications as are shown in Figs. 92,
93, 94; the first of which is a slight advance on the ideal type
represented in Fig. 76, arising in the way described; and the others
of which are actual plants--_Haplomitrium Hookeri_, and _Plagiochila
decipiens_. Thus the higher Archegoniates show us how, along with
an assumption of the upright attitude, there does go on, as we see
there must go on, a separation of the leaf-producing parts from the
root-producing parts; a greater development of that connecting portion
of the successive fronds, by which they are kept in communication with
the roots, and raised above the ground; and a consequent increased
differentiation of such connecting portion from the parts attached
to it. And this lateral bulging of the axis, directly or indirectly
consequent on its functions as a support and a channel, being here
unrestrained by the early-formed fronds folded round it, goes on
without the bursting of these. Hence arises a leading character of
what is called exogenous growth--a growth which is, however, still
habitually accompanied by exfoliation, in flasks, of the outermost
layers, continually being cracked and split by the accumulation of
layers within them. And now if we examine plants of the exogenous type,
we find among them many displaying the stages of this metamorphosis. In
Fig. 95, is shown a form in which the continuity of the axis with the
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