The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 197. And now it remains only to observe, in confirmation of
the foregoing synthesis, that it at once explains for us various
irregularities. When we see leaves sometimes producing leaflets from
their edges or extremities, we recognize in the anomaly a resumption of
an original mode of growth: fronds frequently do this. When we learn
that a flowering plant, as the _Drosera intermedia_, has been known to
develop a young plant from the surface of one of its leaves, we are
at once reminded of the proliferous growths and fructifying organs in
the Liverworts. The occasional production of bulbils by Phænogams,
ceases to be so surprising when we find it to be habitual among the
inferior Acrogens, and when we see that it is but a repetition,
on a higher stage, of that self-detachment which is common among
proliferously-produced fronds. Nor are we any longer without a solution
of that transformation of foliar organs into axial organs, which not
uncommonly takes place. How this last irregularity of development is to
be accounted for, we will here pause a moment to consider. Let us first
glance at our data.
The form of every organism, we have seen, must depend on the structures
of its physiological [or constitutional] units. Any group of such
units will tend to arrange itself into the complete organism, if
uncontrolled and placed in fit conditions. Hence the development of
fertilized germs; and hence the development of those self-detached
cells which characterize some plants. Conversely, physiological units
which form a small group involved in a larger group, and are subject to
all the forces of the larger group, will become subordinate in their
structural arrangements to the larger group--will be co-ordinated into
a part of the major whole, instead of co-ordinating themselves into a
minor whole. This antithesis will be clearly understood on remembering
how, on the one hand, a small detached part of a hydra soon moulds
itself into the shape of an entire hydra; and how, on the other hand,
the cellular mass that buds out in place of a lobster’s lost claw,
gradually assumes the form of a claw--has its parts so moulded as to
complete the structure of the organism: a result which we cannot but
ascribe to the forces which the rest of the organism exerts upon it.
Consequently, among plants, we may expect that whether any portion of
protoplasm moulds itself into the typical form around an axis of its
own, or is moulded into a part subordinate to another axis, will depend
on the relative mass of its physiological units--the accumulation of
them that has taken place before the assumption of any structural
arrangement. A few illustrations will make clear the validity of this
inference. In the compound leaf, Fig. 65, the several lateral growths
_a_, _b_, _c_, _d_, are manifestly homologous; and on comparing a
number of such leaves together, it will be seen that one of these
lateral growths may assume any degree of complexity, according to the
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account