The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§198. The general conclusion to which these various lines of evidence
converge, is, then, that the shoot of a flowering plant is an aggregate
of the third degree of composition. Taking as aggregates of the first
order, those small portions of protoplasm which ordinarily assume
the forms under which they are known as cells; and considering as
aggregates of the second order, those assemblages of such cells which,
in the lower cryptogams, compose the various kinds of thallus; then
that structure, common to the higher cryptogams and to phænogams,
in which we find a series of such groups of cells bound up into a
continuous whole, must be regarded as an aggregate of the third
order. The inference drawn from analysis, and verified by a synthesis
which corresponds in a remarkable manner with the facts, is that
those compound parts which, in Monocotyledons and Dicotyledons are
called axes, have really arisen by integration of such simple parts
as in lower plants are called fronds. Here, on a higher level,
appears to have taken place a repetition of the process already
observed on lower levels. The formation of those small groups of
physiological units which compose the lowest protophytes, is itself
a process of integration; and the consolidation of such groups into
definitely-circumscribed and coherent cells or morphological units, is
a completing of the process. In those coalescences by which many such
cells are joined into threads, and discs, and solid or flattened-out
masses, we see these morphological units aggregating into units of a
compound kind: the different phases of the transition being exemplified
by groups of various sizes, various degrees of cohesion, and various
degrees of definiteness. And now we find evidences of a like process
on a larger scale: the compound groups are again compounded. Moreover,
as before, there are not wanting types of organization by which the
stages of this higher integration are shadowed forth. From fronds that
occasionally produce other fronds from their surfaces, we pass to those
that habitually produce them; from those that do so in an indefinite
manner, to those that do so in a definite manner; and from those that
do so singly, to those that do so doubly and triply through successive
generations of fronds. Even within the limits of a sub-class, we find
gradations between fronds irregularly proliferous, and groups of such
fronds united into a regular series.
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