The Sense of TasteHollingworth, Harry L. (Harry Levi)
Science
The Sense of Taste
Hollingworth, Harry L. (Harry Levi)
Taste
Granting that a dynamic theory as outlined above, with possible
modifications to meet new discoveries, is correct, still another
question requires an answer. In the visual organ affected by ether
vibrations of different rates every theory assumes certain parts of the
mechanism sensitized especially to certain vibration rates. To take the
theory of Hering, for example, there are three photochemical substances,
one decomposed by a relatively slow vibration rate, another by a more
rapid vibration rate, and so on. And in the auditory mechanism there is
the _basilar membrane_, capable of vibration in parts in sympathy with
different rates of vibration of the air. The answer to this question
concerning the taste sense is peculiarly difficult, in that it is
impossible to stimulate individual taste buds, and even difficult to
stimulate a single papilla, which may contain many buds. And then it
must be further borne in mind that each bud contains many nerve endings,
so that what is stimulated in experimental work is really a very complex
portion of the sense organ.
As mentioned in another connection, certain papillæ when stimulated with
various substances respond to but one, e.g., sweet, sour, bitter, or
salt, while others respond to two or three stimuli. Further, it is well
known that certain portions of the tongue when stimulated produce a
predominance of certain kinds of taste sensations. On the other hand, an
examination of the taste buds or of the nerve endings within the taste
buds shows no differences among them. Still, in this respect taste does
not differ from sight. The cones of the retina, which are sensory ends
of vision, show no differences in structure, and it is generally
believed that all of the elementary sensations can be produced by the
stimulation of one single cone. If it be true that the nerve fibrils in
the taste buds themselves receive the stimulus, then any specific
characteristics of the receiving mechanism would have to be looked for
in these nerve endings.
Oehrwall has accounted for the facts of the specific sensitivity of
different parts of the tongue and of different individual papillæ
mentioned above by the assumption that each taste bud has a specific
function and that certain papillæ of the tongue have buds all of one
sort, other of two sorts, and so on. Nagel prefers to modify this view
to the extent that each taste bud is capable of arousing every
elementary sensation, but is adapted to respond best and easiest to a
certain specific stimulus with a certain elementary sensation. This view
is analogous to that of the visual photochemical substances which,
according to the Helmholtz theory, were each sensitive to all wave
lengths of light, but not equally.
Public-domain text, read in full here on John Shaqi.
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