The enormous prey which some snakes are able to swallow is quite
astounding. Anacondas and Pythons, the largest snakes, have been known
to swallow calves and good-sized antelopes with their horns, animals
which, even after being somewhat crushed by constriction, very much
exceed the calibre of the snake. A _Python molurus_ 17 feet long is
reported on good evidence to have swallowed a gravid Axis deer. A
Grass-snake half an inch in diameter can manage a frog or toad three
times that width, and a _Dasypeltis_ of the same size a hen's egg. Such
feats are rendered possible by the mobility of the jaws and
palato-pterygoid arch on the cranium, and the elasticity of the
ligaments by which they are attached (see above, p. 42), as well as by
the mobility of the ribs and the absence of sternal apparatus, together
with the great distensibility of the skin. When a snake proceeds to
dispose of a large prey, which, if it be a mammal or bird, is usually
seized head-first, it pulls itself forward by alternate movements of the
jaws, the maxillary and the mandibular ramus of the one side, and then
of the other, being extended anteriorly and laterally, the snake at the
same time producing an abundant salivation which renders the prey very
slimy. Several repeated alternate movements of the jaws bring the head
of the prey to the gullet, where the muscles and ribs come into play,
and the two sides of the jaws work no longer alternately, but together.
When once in the oesophagus, the prey progresses with much greater
facility, and usually reaches the stomach in a few minutes, whilst the
previous process of deglutition may have lasted half an hour. While this
laborious operation is going on, the breathing of the snake is not
impaired owing to a remarkable contrivance: the trachea can be protruded
in such a manner as to bring its opening outside the mouth.
In cases where the victim is eaten alive, the snake has to contend with
its struggles, but retrogression is rendered impossible by the
backwardly-directed sharp teeth with which the jaws and palate are
beset. A frog is usually caught by one of the hind limbs and swallowed
back-first, the long hind limbs stretching forwards as they fold against
the body; its struggles are often still apparent when it has reached the
oesophagus. Snakes when caught immediately after a meal are in the habit
of disgorging their food, and it sometimes happens that a frog or toad
is thus vomited alive. An instance is known of a naturalist having
captured a Grass-snake and put it in a linen bag. On opening it a short
time after, great was his surprise to find the snake had escaped through
a small hole in the bag, leaving instead a living toad too big to pass
through the hole.
Public-domain text, read in full here on John Shaqi.
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