The Brachiopods have passed into entirely new and more advanced species
in the many advances and retreats of the shores, but the Molluscs show
more interesting progress. The commanding group from the start is that
of the Molluscs which have "kept their head," the Cephalopods, and
their large shells show a most instructive evolution. The first great
representative of the tribe is a straight-shelled Cephalopod, which
becomes "the tyrant and scavenger of the Silurian ocean" (Chamberlin).
Its tapering, conical shell sometimes runs to a length of fifteen
feet, and a diameter of one foot. It would of itself be an important
evolutionary factor in the primitive seas, and might explain more than
one advance in protective armour or retreat into heavy shells. As the
period advances the shell begins to curve, and at last it forms a
close spiral coil. This would be so great an advantage that we are not
surprised to find the coiled type (Goniatites) gain upon and gradually
replace the straight-shelled types (Orthoceratites). The Silurian
ocean swarms with these great shelled Cephalopods, of which the little
Nautilus is now the only survivor.
We will not enlarge on the Sponges and Corals, which are slowly
advancing toward the higher modern types. Two new and very powerful
organisms have appeared, and merit the closest attention. One is the
fish, the remote ancestor of the birds and mammals that will one day
rule the earth. The other may be the ancestor of the fish itself, or it
may be one of the many abortive outcomes and unsuccessful experiments of
the stirring life of the time. And while these new types are themselves
a result of the great and stimulating changes which we have reviewed
and the incessant struggle for food and safety, they in turn enormously
quicken the pace of development. The Dreadnought appears in the
primitive seas; the effect on the fleets of the world of the evolution
of our latest type of battleship gives us a faint idea of the effect, on
all the moving population, of the coming of these monsters of the deep.
The age had not lacked incentives to progress; it now obtains a more
terrible and far-reaching stimulus.
To understand the situation let us see how the battle of land and sea
had proceeded. The Devonian Period had opened with a fresh emergence of
the land, especially in Europe, and great inland seas or lakes were left
in the hollows. The tincture of iron which gives a red colour to our
characteristic Devonian rocks, the Old Red Sandstone, shows us that
the sand was deposited in inland waters. The fish had already been
developed, and the Devonian rocks show it swarming, in great numbers and
variety, in the enclosed seas and round the fringe of the continents.
Public-domain text, read in full here on John Shaqi.
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