The Vertebrate SkeletonReynolds, Sidney H. (Sidney Hugh)
Science
The Vertebrate Skeleton
Reynolds, Sidney H. (Sidney Hugh)
Skeleton
In the HOLOCEPHALI--_Chimaera_, _Hariotta_ and _Callorhynchus_--only
three pairs of teeth or dental plates occur, two pairs in the upper
jaw, one in the lower. These structures persist throughout life and
grow continuously. The upper tooth structures are attached
respectively to the ethmoid or vomerine region of the skull, and to
the palato-pterygoids. The vomerine teeth are small, while those
attached to the mandible and the palato-pterygoid region are large and
bear several roughened ridges adapted for grinding food. The teeth of
the two opposite sides of the jaw meet in a median symphysis. The
teeth of _Chimaera_ are more adapted for cutting, those of
_Callorhynchus_ for crushing. Many extinct forms are known, some of
whose teeth are intermediate in structure between those of _Chimaera_
and _Callorhynchus_.
The teeth of GANOIDS are also extremely variable. Among living forms,
the Holostei are more richly provided with teeth than are any other
fishes, as they may occur on the premaxillae, maxillae, palatines,
pterygoids, parasphenoid, vomers, dentaries, and splenials. Among the
Chondrostei, on the other hand, the adult Acipenseridae are toothless;
small teeth however occur in the larval sturgeon, and in _Polyodon_
many small teeth are found attached merely to the mucous membrane of
the jaws. Many fossil Ganoids have numerous flattened or knob-like
teeth, borne on the maxillae, palatines, vomers and dentaries. Others
have a distinctly heterodont dentition. Thus in _Lepidotus_ the
premaxillae bear chisel-like teeth, while knob-like teeth occur on the
maxillae, palatines and vomers. In _Rhizodus_ all the teeth are
pointed, but while the majority are small a few very large ones are
interspersed.
In TELEOSTEANS, too, the teeth are eminently variable both in form and
mode of arrangement. They may be simple and isolated, or compound, and
may be borne on almost any of the bones bounding the mouth cavity, and
also as in the Pike, on the hyoid and branchial arches. The splenial
however never bears teeth and the pterygoid and parasphenoid only
rarely, thus differing from the arrangement in the Holostei.
The isolated teeth are generally conical in form and are ankylosed to
the bone that bears them. Such teeth are, with a few exceptions such
as _Balistes_, not imbedded in sockets nor replaced vertically.
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