The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammaliaOwen, Richard
Science
The zoology of the voyage of H.M.S. Beagle [vol. 1 of 5] : $b Fossil mammalia
Owen, Richard
Beagle Expedition (1831-1836); Mammals, Fossil; Paleontology -- South America; Zoology
In the fig. 5, Pl. XVII., the fine ivory is represented by the white
striated concentric tract on the grinding surface of the teeth; it is of
a yellowish-white colour in the fossil, and stands out, as an obtuse
ridge, from that surface: both these conditions depend on the large
proportion of the mineral to the animal constituent in this substance of
the tooth. The external layer of the cement presents in the fossil the
same yellowish-brown tint as the bone itself, which it so closely
resembles, both in intimate structure and in chemical composition; the
internal layer next the dense ivory is jet black, indicating the great
proportion of animal matter originally present in this part. The central
pillar of coarse ivory, which, from its more yielding texture, has been
worn down into a hollow at the triturating surface of the tooth, also
presents, as a consequence of the less proportion of the hardening
phosphates, a darker brown colour than the external layer of the cement,
or the bone itself.
The teeth are implanted in very deep sockets; about one-sixth only of
the last molar projects above the alveolus; the proportion of the
exposed part of the tooth increases as they are placed further forwards.
The implanted part of each tooth is simple; preserving the same size and
form as the projecting crown, and presenting a large conical cavity at
the base, indicative of the original persistent pulp, and perpetual
growth of these teeth.
The extent of the whole four alveoli is four inches, eight lines; the
length of the jaw from the angle to the broken end of the symphysis is
seventeen inches and a half;[42] from the figures it will be seen that
only a small proportion of the anterior part of the jaw is lost, so that
we may regard the dentigerous part of the jaw as being limited to about
one-fourth of its entire length; the alveoli being nearly equidistant
from the two extremities. The first and second teeth, counting
backwards, are separated by an interspace of rather more than three
lines; that between the second and third is one line less; the third and
fourth are rather more than a line apart: from the oblique position,
however, of the three hinder teeth the intervals between them appear in
a side view, as in fig. 1, Pl. XIX., to be less than in reality, and the
third and fourth teeth seem to touch each other.
Each tooth has a form and size peculiar to itself, and different from
the rest, but corresponds of course with its fellow on the opposite
side. The same may be observed, but in a less degree, in the teeth of
the Megatherium itself; hence, it is obviously hazardous to found a
generic distinction upon a single tooth, unless, as in the case of the
_Glyptodon_,[43] the modification of form happens to be extremely
well-marked. The whole series of teeth, or their sockets, at least of
one of the jaws, should be known for the purpose of making a
satisfactory comparison with the previously established Edentate genera.
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