Tropical nature, and other essaysWallace, Alfred Russel
History
Tropical nature, and other essays
Wallace, Alfred Russel
Ecology -- Tropics; Natural history -- Tropics; Tropics
Although, as has been shown elsewhere, it may be doubted whether
light directly produces floral colour, there can be no doubt that it
is essential to the growth of vegetation and to the full development
of foliage and of flowers. In the forests all trees, and shrubs,
and creepers struggle upwards to the light, there to expand their
blossoms and ripen their fruit. Hence, perhaps, the abundance of
climbers which make use of their more sturdy companions to reach
this necessary of vegetable life. Yet even on the upper surface
of the forest, fully exposed to the light and heat of the tropical
sun, there is no special development of coloured flowers. When from
some elevated point you can gaze down upon an unbroken expanse
of woody vegetation, it often happens that not a single patch of
bright colour can be discerned. At other times, and especially at
the beginning of the dry season, you may behold scattered at wide
intervals over the mottled-green surface a few masses of yellow,
white, pink, or more rarely of blue colour, indicating the position
of handsome flowering trees.
The well-established relation between coloured flowers and the
need of insects to fertilize them, may perhaps be connected with
the comparative scarcity of the former in the equatorial forests.
The various forms of life are linked together in such mutual
dependence that no one can inordinately increase without bringing
about a corresponding increase or diminution of other forms. The
insects which are best adapted to fertilize flowers cannot probably
increase much beyond definite limits, because in doing so they would
lead to a corresponding increase of insectivorous birds and other
animals which would keep them down. The chief fertilizers--bees
and butterflies--have enemies at every stage of their growth, from
the egg to the perfect insect, and their numbers are, therefore,
limited by causes quite independent of the supply of vegetable
food. It may, therefore, be the case that the numbers of suitable
insects are totally inadequate to the fertilization of the countless
millions of forest-trees over such vast areas as the equatorial
zone presents, and that, in consequence, a large proportion of
the species have become adapted either for self-fertilization or
for cross-fertilization by the agency of the wind. Were there not
some such limitation as this, we should expect that the continued
struggle for existence among the plants of the tropical forests
would have led to the acquisition, by a much larger proportion of
them, of so valuable a character as bright-coloured flowers, this
being almost a necessary preliminary to a participation in the
benefits which have been proved to arise from cross-fertilization
by insect agency.
Public-domain text, read in full here on John Shaqi.
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