A Guide to the Study of Fishes, Volume 1 (of 2)Jordan, David Starr
History
A Guide to the Study of Fishes, Volume 1 (of 2)
Jordan, David Starr
Fishes; Fishes -- Bibliography; Zoology -- History
The homocercal tail is the fan-shaped or symmetrical tail common
among the Teleosts, or bony fishes. In its process of development
the individual tail is first archicercal, then lophocercal, then
diphycercal, then heterocercal, and lastly homocercal. A similar order
is indicated by the sequence of fossil fishes in the rocks, although
some forms of diphycercal tail may be produced by degeneration of the
heterocercal tail, as suggested by Dr. Dollo and Dr. Boulenger, who
divide diphycercal tails into primitive and secondary.
The peculiar tapering tail of the cod, the vertebræ growing
progressively smaller behind, is termed _isocercal_ by Professor Cope.
This form differs little from diphycercal, except in its supposed
derivation from the homocercal type. A similar form is seen in eels.
[Illustration: FIG. 67.--Heterocercal tail of Young Trout, _Salmo
fario_ (Linnæus). (After Parker and Haswell.)]
The term _leptocercal_ has been suggested by Gaudry, 1883, for those
tails in which the vertebral column ends in a point. We may, perhaps,
use it for all such as are attenuate, ending in a long point or whip,
as in the _Macrouridæ_, or grenadiers, the sting-rays, and in various
degenerate members of almost every large group.
The term _gephyrocercal_ is devised by Ryder for fishes in which the
end of the vertebral axis is aborted in the adult, leaving the caudal
elements to be inserted on the end of this axis, thus bridging over
the interval between the vertical fins, as the name (~gephyros~,
bridge; ~kerkos~, tail) is intended to indicate. Such a tail has been
recognized in four genera only, _Mola_, _Ranzania_, _Fierasfer_, and
_Echiodon_, the head-fishes and the pearl-fishes.
[Illustration: FIG. 68.--Isocercal tail of Hake, _Merluccius productus_
(Ayres).]
[Illustration: FIG. 69.--Homocercal tail of a Flounder, _Paralichthys
californicus_.]
The part of the body of the fish which lies behind the vent is known
as the urosome. The urostyle is the name given to a modified bony
structure, originally the end of the notochord, turned upward in most
fishes. The term _opisthure_ is suggested by Ryder for the exserted tip
of the vertebral column, which in some larvæ (_Lepisosteus_) and in
some adult fishes (_Fistularia_, _Chimæra_) projects beyond the caudal
fin. The urosome, or posterior part of the body, must be regarded as
a product of evolution and specialization, its function being largely
that of locomotion. In the theoretically primitive fish there is no
urosome, the alimentary canal, as in the worm, beginning at one end of
the body and terminating at the other.
[Illustration: FIG. 70.--Gephyrocercal tail of _Mola mola_ (Linnæus).
(After Ryder.)]
=Homologies of the Pectoral Limb.=--Dr. Gill has made an elaborate
attempt to work out the homologies of the bones of the pectoral
limb.[4] From his thesis we take the following:
"The following are assumed as premises that will be granted by all
zootomists:
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