A Guide to the Study of Fishes, Volume 2 (of 2)Jordan, David Starr
History
A Guide to the Study of Fishes, Volume 2 (of 2)
Jordan, David Starr
Fishes; Fishes -- Bibliography; Zoology -- History
Several fossil _Halosauridæ_ are described from the Cretaceous of Europe
and Syria, referred to the genera _Echidnocephalus_ and _Enchelurus_.
Boulenger refers the _Lyopomi_ to the suborder _Heteromi_.
CHAPTER VII
THE APODES, OR EEL-LIKE FISHES
=THE Eels.=—We may here break the sequence from the _Isospondyli_ to the
other soft-rayed fishes, to interpolate a large group of uncertain
origin, the series or subclass of eels.
The mass of apodal or eel-like fishes has been usually regarded as
constituting a single order, the Apodes (ἄ, without; ποῦς, foot). The
group as a whole is characterized by the almost universal separation of
the shoulder-girdle from the skull, by the absence of the mesocoracoid
arch on the shoulder-girdle, by the presence of more than five pectoral
actinosts, as in the Ganoid fishes, by the presence of great numbers of
undifferentiated vertebræ, giving the body a snake-like form, by the
absence in all living forms of the ventral fins, and, in all living
forms, by the absence of a separate caudal fin. These structures
indicate a low organization. Some of them are certainly results of
degeneration, and others are perhaps indications of primitive
simplicity. Within the limits of the group are seen other features of
degeneration, notably shown in the progressive loss of the bones of the
upper jaw and the membrane-bones of the head and the degradation of the
various fins. The symplectic bone is wanting, the notochord is more or
less persistent, the vertebral centra always complete constricted
cylinders, none coalesced. But, notwithstanding great differences in
these regards, the forms have been usually left in a single order, the
more degraded forms being regarded as descended from the types which
approach nearest to the ordinary fishes. From this view Professor Cope
dissents. He recognizes several orders of eels, claiming that we should
not unite all these various fishes into a single order on account of the
eel-like form. If we do so, we should place in another order those with
the fish-like form. It is probable, though not absolutely certain, that
the _Apodes_ are related to each other. The loss among them, first, of
the connection of the post-temporal with the skull; second, of the
separate caudal fin and its hypural support; third, of the distinct
maxillary and premaxillary; and fourth, of the pectoral fins, must be
regarded as successive phases of a general line of degradation. The
large number of actinosts, the persistence of the notochord, the absence
of spines, and the large numbers of vertebræ seem to be traits of
primitive simplicity. Special lines of degeneration are further shown by
deep-sea forms. What the origin of the _Apodes_ may have been is not
known with any certainty. They are soft-rayed fishes, with the
air-bladder connected by a tube with the œsophagus, and with the
anterior vertebræ not modified. In so far they agree with the
_Isospondyli_.
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