A Guide to the Study of Fishes, Volume 2 (of 2)Jordan, David Starr
History
A Guide to the Study of Fishes, Volume 2 (of 2)
Jordan, David Starr
Fishes; Fishes -- Bibliography; Zoology -- History
=SUBORDER Hemibranchii.=—Still another transitional group, the
_Hemibranchii_, is composed of spiny-rayed fishes with abdominal
ventrals. In this suborder there are other points of divergence, though
none of high importance. In these fishes the bones of the
shoulder-girdle are somewhat distorted, the supraclavicle reduced or
wanting, and the gill structures somewhat degenerate. The presence of
bones called interclavicles or infraclavicles, below and behind the
clavicle, has been supposed to characterize the order of _Hemibranchii_.
But this character has very slight importance. In two families,
_Macrorhamphosidæ_ and _Centriscidæ_, the interclavicles are absent
altogether. In the _Fistulariidæ_ they are very large. According to the
studies of Mr. Edwin C. Starks, the bone in question is not a true
infraclavicle. It is not identical with the infraclavicle of the
Ganoids, but it is only a backward extension of the hypocoracoid, there
being no suture between the two bones. In those species which have bony
plates instead of scales, this bone has a deposit of bony substance or
ganoid enamel at the surface. This gives it an apparent prominence as
compared with other bones of the skeleton, but it has no great taxonomic
importance. Dr. Hay unites the suborders _Hemibranchii_,
_Lophobranchii_, and _Hypostomides_ to form the order _Phthinobranchii_
(φθινάς, waning; βράγχος, gill), characterized by the reduction of the
gill-arches. These forms are really nearly related, but their affinities
with the _Percesoces_ are so close that it may not be necessary to form
a distinct order of the combined group. Boulenger unites the
_Hemibranchii_ with _Lampris_ to form a group, _Catosteomi_,
characterized by the development of infraclavicles; but we cannot see
that _Lampris_ bears any affinity to the sticklebacks, or that the
presence of infraclavicle has any high significance, nor is it the
supposed infraclavicle of _Lampris_ homologous with that of the
_Hemibranchii_. The dorsal fin in the _Hemibranchii_ has more or less
developed spines; spines are also present in the ventral fins. The lower
pharyngeals are separated; there is no air-duct. The mouth is small and
the bones of the snout are often much produced. The preopercle and
symplectic are distinct. The group is doubtless derived from some
transitional spiny-rayed type allied to the _Percesoces_. The
Lophobranchs, another supposed order, represent simply a still further
phase of degradation of gills and ventral fins. Dr. Gill separates these
two groups as distinct orders and places them, as aberrant offshoots,
near the end of his series of bony fishes. We prefer to leave them with
the other transitional forms, not regarding their traits of divergence
as of any great importance in the systematic arrangement of families.
Public-domain text, read in full here on John Shaqi.
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