Order 6. =Agaricaceæ= (=Mushrooms=, =Toadstools=). _The hymenophore
consists_ of knife-like plates (_lamellæ_, _gills_), which are situated
on the under side of the umbrella-like pileus of the fruit-body, and
radiate from the central stalk. Those which are first formed extend
from the edge of the pileus to the stalk; those formed later reach only
a longer or shorter portion of this distance, according to their age.
In structure the lamellæ (Fig. 174) consist of a central mass of hyphæ,
the _trama_, continuous with the hyphæ of the pileus; these terminate
in a layer of shorter cells, the _subhymenial layer_, immediately
beneath the hymenium which is composed of basidia and paraphyses. In a
few species, but not in the majority, the lamellæ are branched, and in
some they are decurrent. A few have the stalk placed excentrically, or
it may be entirely absent.
[Illustration: FIG. 172.--Development of _Psalliota campestris_:
_a_, _b_, _c_, _d_ show the various stages of the development
of the fruit-bodies and the mycelium (_m_) (nat. size); _e_ the
fruit-body in a somewhat later stage, slightly magnified; _f_
longitudinal section of _e_; _n_ first formation of the hymenium;
_g_ longitudinal section of a more advanced fruit-body (nat.
size); _n_ the hymenium; _o_ velum partiale (see Fig. 133.)]
In the early stages of its development the fruit-body is more or
less enclosed in a hyphal tissue--the “veil” (_velum universale_, or
_volva_). The veil at first completely encloses the young fruit-body,
but is afterwards ruptured as the latter grows, part remaining at the
base of the stalk as the “sheath” (_annulus inferus_), and part on the
pileus as scales or warts. In the “Fly Mushroom” (_Amanita muscaria_)
the remains of the veil are especially conspicuous as white patches
on the bright red ground of the upper surface of the pileus, and as
a sheath at the base of the stalk (Fig. 178 _v._). Another veil--the
_velum partiale_--a hyphal tissue (Figs. 178 _a_; 173) stretches from
the edge of the pileus to the stalk, and encloses the lamellæ. This
veil is ruptured as the pileus expands, a portion attached to the stalk
remaining as the “upper ring” (_annulus superus_) (Figs. 173, 178 _a_),
or a part attached to the pileus hanging down as a fringe round its
edge.--Some genera have no veil, the under side of the pileus being
exposed from the first (_gymnocarpic_ Agaricaceæ). Those which have
a veil (_hemiangiocarpic_ A.) afford a transition to the angiocarpic
Gasteromycetes.
[Illustration: FIG. 173.--The cultivated Mushroom (_Psalliota
campestris_).]
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