The internal structure of the sexual generation is very simple. The
leaves in nearly all cases are formed of a single-layered plate of
cells; in the Leafy-Mosses, however, a midrib is very often formed, and
sometimes, also, marginal veins; and along these lines the leaves are
several layers of cells in thickness. The stem is constructed of cells
longitudinally elongated, the external ones of which are narrower and
sometimes have thicker walls than the more central ones. _Vessels are
not found_, but in several Mosses there is in the centre of the stem
a conducting strand of narrow, longitudinal cells, which represents
the vascular bundle in its first stage of development. This strand
contains elements for conveying water as well as sieve-tubes. Stomata
are entirely wanting in the sexual generation of the Leafy-Mosses; they
are found in a few Liverworts (_Marchantia_), but their structure is
not the same as in the higher plants.
VEGETATIVE REPRODUCTION takes place by gemmæ or buds which arise on
the protenema, the rhizoids, the thallus, or the shoots, and become
detached from the mother-plant; or else the protonema and the older
parts of the plant simply die off, and their branches thus become
independent plants. This well-developed vegetative reproduction
explains why so many Mosses grow gregariously. In certain Marchantiaceæ
special cupules, in which gemmæ are developed, are found on the surface
of the thallus (Fig. 197 _A_, _s-s_). Again, protonema may also arise
from the leaves, and thus the leaves may act as reproductive bodies.
Certain Mosses nearly always reproduce vegetatively, and in these
species the oospheres are seldom fertilised.
[Illustration: FIG. 187.--_Marchantia polymorpha_: _a_ mature
antheridium.]
[Illustration: FIG. 188.--Spermatozoids.]
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