Five species of _Lycopodium_ are found in Great Britain. _L.
clavatum_ and _L. selago_ are common in mountainous districts.
_L. annotinum_ is common in the Highlands of Scotland. The other
genus of the order is _Phylloglossum_, with one species, _P.
drummondi_ (Australia, Tasmania, and New Zealand), a small plant
only a few centimetres high, with two tubers, and about eleven
linear leaves at the base of the stem which is terminated by a
cone of sporophylls.--FOSSIL Lycopodiaceæ in the Carboniferous
period.
OFFICINAL: “Lycopodium,” the spores of _L. clavatum_.
Family 2. =Psilotaceæ=. The sporangia are placed on the apex of
short, two-leaved stems, as 2–3, seldom four, small capsules.
Small herbs, with angular stems; leaves small, simple, and one
nerved. Only four species.--_Psilotum_ (Madagascar, Moluccas,
Sandwich Islands, etc.) is destitute of roots, their place being
supplied by special underground stems which bear a few modified
leaves, very much reduced, especially when buried deeply in the
soil. Three species.--_Tmesipteris_ (Australia), one species.
Sub-Class 2. =Selaginelleæ= (HETEROSPOROUS Lycopodinæ).
Micro-and macrospores. The prothallia are very much reduced, especially
the male; the female does not leave the spore. The leaves are ligulate.
[Illustration: FIG. 233.--Germination of the microspores of
_Selaginella_: _A_ the spore rendered transparent, seen from
above. In the interior is seen the prothallium (_f_), and the
first divisions of the antheridium (_a_, _b_, _c_, _d_); in _B_
the spore-wall is removed and all the spermatozoid-mother-cells
formed; in _C_, the microspore has opened and the spermatozoids
and the mother-cells are escaping together.]
=The sexual generation.= In the ~MICROSPORES~ are formed: (1) a very
small “vegetative” cell, representing the vegetative part of the
prothallium (_f_ in Fig. 233 _A, B_), and (2) a cell many times larger
and which divides into a number (4–8) of primordial cells, each of
which divides into four spermatozoid-mother-cells, though all of these
may not develope spermatozoids. On germination, when the spore-wall is
ruptured, the spermatozoids and spermatozoid-mother-cells are ejected
into the water.
The ~SPERMATOZOIDS~ in _Selaginella_ are elongated and club-shaped,
with two cilia (Fig. 234); but in _Isoëtes lacustris_ they are
spirally-twisted threads which differ from all other spermatozoids by
having a bunch of cilia _at each end_; the other species of _Isoëtes_
have cilia only at the anterior end.
The MACROSPORES. Shortly after the macrospores have been set free,
or in _Selaginella_, while still enclosed in the sporangium of the
mother-plant, they germinate and soon become filled with the cellular
tissue of the prothallium, and even in _Selaginella_ the archegonium
begins to be formed before the rupture of the spore-cell-wall has
commenced (Fig. 235 _A_).
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