of its nourishment during germination. These are termed albuminous
(endospermous) seeds, in distinction to the ex-albuminous, or those in
which the nutritive-tissue is stored in the embryo itself, before it is
completely developed, and used for its sustenance.
In addition to the changes which fertilisation produces in the ovule
itself, it also gives the impetus to a series of changes in the
entire shoot which bears the ovule. The perianth, stamens, and style,
generally wither, because the part they play is at an end; the wall
of the ovary grows and becomes the wall of the fruit (pericarp).
The entire gynœcium of a flower, transformed as a consequence of
fertilisation, is termed a _fruit_. It consists of two parts, the
_pericarp_ and the _seeds_, and according to the nature of the
pericarp, the fruit is termed a capsule, nut, berry, or drupe.
The chief characteristic of the Phanerogams does not lie in the
formation of the flower (although they may quite properly be termed
“Flowering-plants”), because Equisetums and Lycopods have reproductive
shoots as highly differentiated as those of certain Gymnosperms and
other Phanerogams. As regards the SEXUAL GENERATION the characteristics
are found:--(1) in its great reduction; (2) in the transmission of the
microspore (pollen-grain) to the macrosporangium, and its germination,
with the formation of a _pollen-tube_ (antheridium), the protoplasm of
which is not differentiated into spermatozoids; (3) in the fact that
the macrospore (embryo-sac) never leaves its sporangium (nucellus);
and further in the Angiosperms, (4) in the peculiar development of the
nutritive-tissue in two parts; and (5) in the great reduction of the
archegonium.
As regards the ASEXUAL GENERATION the characteristic feature is that
this generation is formed whilst the sporangium is still attached to
the mother-plant, and for a long time is nourished by it; and that
after the sporangium has become detached from the mother-plant, it
spends a longer or shorter resting period as the embryo in the seed
(enveloped by the testa), and does not make its appearance until
the “germination” of the seed. In addition the shoot which bears
sporangia undergoes greater modification than in the case of the
Flowerless-plants.
The Phanerogams are separated into two Divisions as follows:--
Division 4. =Gymnospermæ.= The ovules, as well as the seeds, are borne
_naked_ on the surface of _open carpels_, or on the apex of a stem
(ovary wanting). The pollen-grains are conveyed by the wind to the
ovules, and caught by drops of mucilage, secreted by the micropyle.
A “stigma” is _wanting_. The entire _female prothallium_ (_the
endosperm_), which serves for the nourishment of the embryo, is _formed
before fertilisation_. The archegonia are _embedded in the upper part
of the prothallium. The pollen-grains are “multicellular,” i.e._ there
is always in their interior a distinct prothallium, formed by 1–3
cells, and a larger cell which gives rise to the pollen-tube.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account