POLLINATION is accomplished by means of the wind. At the period
of pollination the leaves are always so widely separated from one
another, that the ovules can catch the pollen-grains carried to
them by the wind; this is often effected by the mucilaginous drops
which appear at the micropyle, and by the evaporation of which the
pollen-grains are brought in contact with the nucellus. The entire cone
grows considerably as soon as fertilisation has taken place, and the
cone-scales in Pinoideæ close together so that the seeds while maturing
are enclosed, and it is not until the seeds are ready for distribution
that the cone-scales again become separated. In the Pinoideæ, the fully
developed ovuliferous scales are hard and woody; and in this condition
the collection of female flowers is termed a _cone_. In the Taxoideæ,
true cones are the exception. 2–15 cotyledons are present, arranged in
a whorl.
The characteristic feature of this class is the abundance of _resin_,
which is to be found in isolated cells (especially in the cortex),
partly in intercellular glands or passages (both in the cortex and
wood). _Taxus_ is the only genus which has no resin.
There are about 350 species, mostly from the Northern Temperate
zone (especially North America and Siberia), where they grow
gregariously and form the most northern forests. The Juniper,
Scotch Fir, and Yew are natives of Great Britain.
This class may be divided into two families:--
1. =Taxoideæ.= The ovules have either one integument, the external
part of which is fleshy, and the internal hard and stone-like; or two
integuments, of which the external is the fleshy and coloured “aril.”
“Ovuliferous scales” are wanting. The cones are never woody, but are
generally succulent, the bracts become fleshy, or cones usually are not
developed. The seeds project more or less freely beyond the bracts.
2. =Pinoideæ.= The ovules have two integuments, the external one of
which is leaf-like and becomes developed as the “ovuliferous scale”;
if there are several of these in each flower they unite and form a
“symphyllodium.” This may remain free or unite with the bract. The
cones are most frequently woody, rarely succulent. The seeds are hidden
among the cone-scales.
Family 1. =Taxoideæ.=
This family, considered to be most nearly related to the Cycadeæ, also
made its appearance at a very early period. There is only one order.
Order. =Taxaceæ.= The characters have been given above.
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