THE STEM is frequently a corm, bulb, or other variety of underground
stem, as the majority of the Monocotyledons are perennial, herbaceous
plants; it has scattered, closed vascular bundles (Fig. 276), and no
cambium by which a continuous thickening may take place. The stem of
the Palms, however, attains a very considerable thickness, which is due
to the meristem of its growing-point continually increasing in diameter
for a lengthened period (often for many years), until it has reached a
certain size. In this condition the growing-point has the form of an
inverted cone, and it is only when this cone has attained its requisite
size that the formation of a vertical cylindrical stem commences.
Certain tree-like Liliaceæ, as _Dracæna_, _Aloe_, etc., have a
continuous increase in thickness; this is due to a meristematic layer,
which arises in the cortex, outside the original vascular bundles,
which were formed at the growing-point of the stem. This meristem
continues to form thick-walled parenchyma and new, scattered vascular
bundles. The primary vascular bundles, in the Palms and others, run
in a curved line from their entrance into the stem at the base of
the leaf, towards the centre of the stem, and then bend outwards and
proceed downwards in a direction more parallel to the sides of the stem
(Fig. 277). The bundles formed later, in those stems which increase in
thickness, are not continued into the leaves.
THE BRANCHING as a rule is very slight, the axillary buds of the
majority of the leaves never attaining development, _e.g._ in the
Palms, bulbous plants and others. As the cotyledon arises singly,
the succeeding leaves also must be scattered, but they are frequently
arranged in two rows (Grasses, Iris, etc). _The first leaf borne on a
branch_ (the “Fore-leaf,”[24]--the bracteole, if on a floral shoot)
has generally, in the Monocotyledons, a characteristic form and
position, being situated on the posterior side of its own shoot, and
hence turned towards the main axis; it is sometimes provided with two
laterally-placed keels (Figs. 279 _f_, 290 _øi_), but the midrib is
often absent. It arises in some cases from two primordia, which at the
beginning are quite distinct, and thus has been regarded as formed by
two leaves. It is, however, only one leaf, a fact which is evident from
several circumstances, one being that it never supports more than one
shoot, and this stands in the median plane (Fig. 279).
[Illustration: FIG. 276.--Transverse section of the stem of a
Palm: _v v_ is the wood portion, _b b_ the bast portion of
the vascular bundled.]
[Illustration: FIG. 277.--Diagrammatic representation of the
course of the vascular bundles, from the stem into the leaves in
a Monocotyledon.]
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