_Zostera_ (Grass-wrack) is an entirely submerged, marine plant
with creeping rhizome (with displacement of buds) and strap-shaped
leaves. The flowering shoots are sympodia with displacement of the
axes (Fig. 281). The inflorescence is a peculiar, flatly-compressed
spike, on _one_ side of which the flowers are borne (Fig. 281). ~This
inflorescence may be considered, no doubt correctly, to be derived
from the symmetrical spike of _Potamogeton_ by strongly dorsiventral
development, and by a strong suppression of the floral parts taking
place simultaneously. Two rows of flowers are developed, but of these
one is so pressed into the other that apparently only one is present.~
Each flower consists of only 1 stamen and 1 carpel situated at the same
height (Fig. 281); the unilocular ovary encloses 1 pendulous ovule
and bears a bifid style. As regards the perianth (?) one leaf may be
present (_Z. nana_, Fig. 281 _D_). The pollen-grains are filamentous.
Pollination takes place under water. ~_Posidonia_ and _Cymodocea_ are
allied to these. About 70 species.~
[Illustration: FIG. 281.--_Zostera._ A Diagram of the
branching of the floral shoots: _I_, _II_ ... are the successive
shoot-generations, every other one being shaded; _g_{1}_
_g_{2}_ ... fore-leaves; _sp_{1} sp_{2}_ ... spathes for the
successive spikes. Each shoot is united for some distance with
the parent axis (indicated by the half-shaded internodes). Each
shoot commences with a fore-leaf turning towards the parent
axis, _g_; succeeding this is the spathe, _sp_; and then the
inflorescence. The fore-leaf supports a new lateral shoot.
_B_ Diagram of a shoot, _II_, which is borne laterally in the
axil of the fore-leaf _g_{1}_, on the shoot _I_, _g_{2}_ its
fore-leaf; _sp_{2}_ its spathe; _sti_ squamulæ intravaginales.
_II_ Is the spadix with stamens and carpels; _b_ a perianth-leaf
(or connective expansion, similar to those which occur in
_Potamogeton_). _C_ The upper portion of a young spadix with
development of flowers. _D_ Part of a spadix with 2 flowers; the
parts which theoretically belong to one another are connected by
a dotted line.]
Order 3. =Aponogetonaceæ.= Aquatic plants with tuberous stem. They
have a single, petaloid perianth (3–2–1–leaved), most frequently 6
stamens and 3(-6) carpels. Straight embryo.--About 15 species (Africa,
Madagascar, Tropical Asia and Australia).--_Aponogeton distachyos_ and
_A._ (_Ouvirandra_) _fenestralis_ are grown in conservatories; the
latter has lattice-like, perforated leaves.
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