singly and spirally, or without any definite order, directly
from the main axis, with the production of a _cylindrical_
inflorescence bearing “spikes” _on all sides_, that is, a
“_spike-like panicle_.”--Many inflorescences are somewhat
dorsiventral. The _flower_ is rarely unisexual (_Zea mais_) or
barren. Considerable difficulty is experienced in reducing the
Grass-flower to the ordinary 3-merous Monocotyledonous type.
Some authorities consider the _lodicules_, which are present
in all Grasses but absent in the Cyperaceæ, to be homologous
with a perianth. According to a more recent theory they are
bracteoles, and hence the Gramineæ, like many of the Juncaceæ,
have 2–3 bracteoles placed in two rows in the median plane. If
this theory be correct, the _flower is naked_. The lodicules
expand quickly and cause the opening of the flower (_i.e._ the
two pales become separated from each other). Generally only 3
_stamens_ belonging to the outer whorl are present (Fig. 290),
as in _Iris_ (Fig. 279), certain Juncaceæ and Cyperaceæ (Fig.
284), but in some, such as the Rice and certain species of
Bamboos, all 6 are found. _Pariana_ has more than 6. Only 1 of
the _carpels_ is present, namely, the anterior (of those in Fig.
284), so that the ventral suture and the place of attachment
of the ovule are situated at the back of the ovary. The number
of styles does not correspond with the number of carpels, and
the styles may therefore be supposed to arise from the edges
of the leaf to the right and left--a position which is not
without analogy. In addition, a stylar projection is sometimes
found on the anterior side and in the median line (_e.g._ in
_Phragmites_), and the solitary style in _Nardus_ has exactly
this position; a similar arrangement is found in some species of
_Bambusa_ which have only one style; other species of _Bambusa_
have three styles. A tripartite style is found in _Pharus_.
[The Grass-flower may be reduced to the ordinary
Monocotyledonous type thus:--The outer pale is the bract of the
flower since it bears in its axil the floral shoot; the inner
pale occupies the customary position of the bracteole. The fact
that it is binerved can be explained by its having been pressed
against the main-axis during development. Similar binerved
bracteoles are found in _Iris_ (Fig. 279). These bracteoles in
both Grass and Iris arise from single primordia, and are not
produced by the coalescence of two leaves. The lodicules are the
only parts of the perianth remaining, the outer whorl having
been suppressed, and also the posterior leaf of the inner whorl;
a posterior lodicule, however, is found in the Rice and some
species of Bamboo. The outer whorl of stamens is usually absent,
though this again is present in the Rice and Bamboo. The three
carpels are reduced to one with two or sometimes three stigmas.]
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