_unisexual_ (monœcious or diœcious) with the usual diagram (Fig.
278); the perianth is inconspicuous, green or yellow, persistent, and
more or less leathery or fleshy. 6, rarely 3 or many stamens. The 3
carpels remain either _distinct_ or form one, generally 3-locular,
ovary. The style is short. There is _one ovule in each carpel_. Often
during ripening 2 carpels with their ovules are aborted. The fruit is
a _berry_, _drupe_ or _nut_, generally one-seeded, with a large horny
or bony endosperm with hard thick-walled cells (_e.g._ Date-palm). In
some (_e.g._ Cocoanut) it is thin-walled, soft, and oily; in several
“ruminate.”
When _germination_ commences in the Cocoanut, Date, etc., the
apex of the cotyledon remains in the seed and developes into
a spongy mass to withdraw the endosperm; in the Cocoanut it
attains a considerable size (Fig. 299 _C_) and assumes the
form of the fruit. The endosperm in the Cocoanut is hollow and
the interior is filled with “milk.” In the Date-palm and the
Vegetable-ivory (_Phytelephas_) the cell-walls of the hard
endosperm serve as reserve material.
=1.= PHŒNICEÆ. _Phœnix_ (Date-palm) has pinnate leaves with channeled
leaflets and diœcious flowers with 8 free carpels, of which usually
only one developes into a berry with membranous endocarp; the large
seed has a deep furrow on the inner side, and horny endosperm.
[Illustration: FIG. 297.--Inflorescence of a Palm with spathe. At
the top ♂-, at the base ♀-flowers.]
[Illustration: FIG. 298.--_Livistona australis._]
=2.= SABALEÆ. These have fan-like leaves with channeled segments;
flowers ☿ or polygamous, rarely diœcious, with 3 separate or only
slightly united carpels, all of which are sometimes developed
into fruits (berry or drupe, with thin stone).--_Chamærops_, the
Dwarf-palm. The pericarp is externally fleshy, internally more
fibrous, and provided with a membranous inner layer. The endosperm is
ruminate (that is, the testa is several times deeply folded into the
endosperm).--_Sabal_, _Copernicia_, _Livistona_ (Fig. 298), _Thrinax_,
_Corypha_, _Brahea_, and others.
[Illustration: FIG. 299.--_A_ Longitudinal section of a Cocoanut
(diminished), the inner layer only (the stone) not being divided
_B_ End view of the stone, showing the sutures for the 3 carpels
(_a_), and the 3 germ-pores; the embryo emerges from the lowest
one when germination begins. _C_ Germinating; inside the stone is
seen the hollow endosperm and the enlarging cotyledon.]
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