Order 3. =Pandanaceæ= (Screw-pines) is another small order,
forming a transition to the Araceæ. The woody, (apparently)
dichotomous stem is supported by a large number of aerial roots,
which sometimes entirely support it when the lower portion of
the stem has decayed. The leaves are closely crowded together,
and arranged on the branches in three rows, which are often
obliquely displaced, with the formation of three spiral lines;
they are, as in the Bromeliaceæ, amplexicaul, long, linear,
the edge and lower midrib often provided with thorns. The
♂-flowers are borne in branched, the ♀ in unbranched spadices
or capitula, which resemble those of _Sparganium_, but have no
floral-leaves. Perianth absent. The drupes or berries unite into
multiple fruits.--About 80 species in the islands of the Indian
Ocean.--_Pandanus_, _Freycinetia_.--Fossils perhaps in the chalk
of the Harz.
Order 4. =Typhaceæ.= The flowers are unisexual, monœcious, and borne
on a cylindrical spike or globose capitulum; ♂ inflorescences above,
the ♀ below. The perianth consists of a definite number of scales
(_Sparganium_), or in its place numerous irregularly-arranged hairs are
found (_Typha_); in the ♂-flower there are generally three stamens; the
gynœceum is formed of 1–2 carpels with 1 prolonged style; 1 pendulous
ovule. The seeds are furnished with a seed-cover, which is cast off on
germination.--The few species (about 20) which belong to this order are
marsh plants with creeping rhizome (and hence grow in clusters); the
leaves on the aerial shoots are borne in two rows, entire, very long
and linear.
_Sparganium_ (Bur-reed). The flowers are borne in globose capitula;
the perianth distinct, generally consisting of 3 small scales; pistil
bicarpellate. Drupe, dry and woody. ~The stalk of the lower ♀ capitula
is sometimes united with the main axis, and consequently the capitula
are situated high above their subtending-leaf.~
_Typha_ (Bulrush, Reed-mace) has a long, cylindrical, brown spike,
the lower portion bearing ♀-flowers, and the upper ♂-flowers, which
is divided into joints by alternate leaves. The ♀-flowers have 1
carpel. The perianth is wanting, represented by a number of fine,
irregularly-placed hairs; pistil unicarpellate. Fruit a nut.
The two genera, according to some, are related to the 2nd order.
In both genera native species are found. The pollination is
effected by the wind, and consequently the anthers project
considerably, and the stigma is large and hairy. _Typha_ is
protandrous, _Sparganium_ protogynous. The small, fine hairs
surrounding the nut of _Typha_ assist in its distribution by the
wind.--Fossil _Typhas_ in the Tertiary.
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