HYPOGYNOUS flowers: Colchicaceæ, Liliaceæ, Convallariaceæ,
Bromeliaceæ (in part).
EPIGYNOUS flowers: Amaryllidaceæ, Iridaceæ, Bromeliaceæ (in
part), Dioscoreaceæ.
Order 1. =Colchicaceæ.= The flower (Fig. 304 _A_) is ☿, regular,
_hypogynous_, trimerous in all five whorls (6 _stamens_); anthers
usually _extrorse_. Gynœceum with 3 _free styles_ (_A, D_); fruit a
_capsule with septicidal dehiscence_ (_E_); embryo very small (_F_).
The underground stem is generally a corm or rhizome, seldom a bulb.
=A.= VERATREÆ.--_Veratrum_; perennial herbs, stem tall with long
internodes and broad, folded leaves; the flowers andromonœcious,
with free, widely opening perianth-leaves (Fig. 304 _A_), and
globular anthers; inflorescence a panicle.--_Zygadenus, Melanthium,
Schœnocaulon, Uvularia, Tricyrtis_.
=B.= TOFIELDIEÆ.--_Narthecium_ and _Tofieldia_ have leaves alternate
(arranged in two rows), sword-like and borne in rosettes; racemes
or spikes. _Narthecium_ forms an exception to the order by having
a simple style and fruit with loculicidal dehiscence; _Tofieldia_
by the introrse anthers. In this they are related to the Liliaceæ.
_Narthecium_ has poisonous properties, like many other Colchicaceæ.
[Illustration: FIG. 304.--_Veratrum_: _A_ flower; _B_ stamen;
_C_ transverse section of ovary; _D_ gynœceum, with one carpel
bisected longitudinally, and the third removed; _E_ fruit after
dehiscence; _F_ longitudinal section of a seed.]
=C.= COLCHICEÆ.--_Colchicum_ (Autumn Crocus); perennial herbs, with a
long, _funnel-shaped, gamophyllous perianth_, and introrse anthers. The
flowers of _C. autumnale_ spring up immediately from the underground
stem, which is in reality a _corm_ formed of one internode. ~_Colchicum
autumnale_ flowers in autumn without leaves; in spring the radical
foliage-leaves appear simultaneously with the fruit. The flower is
protogynous, and is pollinated by insects (humble-bees, etc.) which
seek the honey secreted by the free part of the stamen a little way
down the tube. The length of the tube protects the fruit, and not,
as in other cases, the nectary.--_Bulbocodium_ and _Merendera_ have
unguiculate perianth-leaves, free, but closing together like a tube.~
175 species; chiefly in North America and South Africa.
_Tofieldia_ is an Arctic plant. The order is rich in pungent,
poisonous alkaloids (veratrin, colchicin, etc.). OFFICINAL;
the seeds of _Colchicum autumnale_ (Europe) and _Schœnocaulon
officinale_ (Mexico), and the rhizome of _Veratrum album_
(mountains of Central Europe).
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