Order 8. The =Iridaceæ= have _epigynous_, hermaphrodite flowers with
petaloid perianth as in the Amaryllidaceæ, but the _interior whorl of
stamens is entirely suppressed_, and the 3 developed _outer_ stamens
have _extrorse_ anthers (Fig. 279); there is 1 style with 3 large,
generally _more or less leaf-like branches bearing the stigmas_.
Ovary and capsule as in the Amaryllidaceæ and Liliaceæ.--Perennial
herbs; _bulbs are rarely found_, but horizontal rhizomes, corms, etc.,
take their place. The leaves are (except _Crocus_) as in the _Iris_,
_two-rowed_, _equitant_ and _sword-like_. Flowers or inflorescences
terminal.
The _Iris_ (Flag) has a horizontal rhizome. The flowers are borne in
the leaf-axils in fan-like inflorescences (rhipidium). The branches of
the style are large and _petaloid_; on their under surface may be seen
a small projecting shelf (Fig. 312 _a_) having on its upper surface the
stigmatic hairs. Beneath the branches of the style are 3 well protected
stamens, and immediately outside these the external perianth-leaves.
~The honey is secreted in the perianth-tube, and the insects,
endeavouring to obtain it through the narrow passages at the base of
the stamens, settle upon the outer perianth-leaves, which are bent
backwards and often very hairy along their central line. The insects
then rub their backs on the anthers just above them, beneath the
branches of the style; they readily deposit the pollen on the stigma
of another flower as they enter it, but cannot do so in withdrawing,
since the stigma is pushed back, and self-fertilisation is thus
avoided. The stylar branches lie close to the outer perianth-leaves,
which are just beneath them, or separated by a distance of only 6–10
mm.; the first form of flower is adapted for _Rhingia rostrata_, the
latter for bees~.--_Crocus_ has vertical, _tuberous_, underground stems
surrounded by the leaf-sheaths (corms), and terminal flowers; the
linear leaves _are not equitant_, but have two longitudinal furrows
on the under side. The perianth is gamophyllous and funnel-shaped.
The stylar branches (stigmas) are fleshy, _rolled together in the
shape of a horn_, and split along the edge.--_Gladiolus_ has corms
like the _Crocus_; spikes with slightly zygomorphic, almost bilabiate
flowers, most frequently turning to one side. Position of the leaves
as in the Iris.--~_Diplarrhena_ has 2 fertile and 1 barren stamen;
_Hermodactylus_ has a unilocular ovary with 3 parietal placentæ.
_Cypella_ and _Tigridia_ have bulbs.~
[Illustration: FIG. 312.--_Iris pseudacorus._ One external and
two internal perianth-leaves, and one of the stylar-branches
have been removed, _y_ The outer, _i_ the inner perianth-leaves;
_g_ stylar-branch; _a_ stigma; _s_ anther. The ovary is seen in
longitudinal section.]
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