_Corylus_ (Hazel-nut, Fig. 329). The long, cylindrical ♂-catkins pass
the winter naked, 2–3 together, on short branches. The very small
♀-catkins are enclosed in buds, in which they pass the winter; these
buds are situated in the axils of the fallen foliage-leaves, and it
is only by their larger size that they may be distinguished from the
ordinary foliage-buds. In spring the ♀-catkins are easily recognised
by their red, projecting stigmas (Fig. 329 _A_). The cupule--the
“husk”--is tubular, fringed, and envelopes the nut. ~The leaves are
alternate and unsymmetrical, the external side being larger than
the internal; this is connected with the vernation, the blade being
conduplicate in the bud; the stipules are deciduous. The bud-scales
are formed of stipules, the most internal having a leaf-blade attached
to them which is suppressed in the external ones. The cotyledons remain
underground on germination.~
[Illustration: FIG. 329.--_Corylus avellana_: _A_ branch at
the time of flowering with ♂-and ♀-catkins; _B_ ♂-flower with
subtending-leaf (bract) and two bracteoles; _C_ the same without
the anthers; _D_ view of interior of ♀-dichasium shortly after
fertilisation; _E_ young fruit with cupule; _F_ similar one with
the cupule opened; _G_ mature ♀-fruits; _H_ nut.]
_Carpinus_ (_C. betulus_, Hornbeam). The ♂-and ♀-catkins do not appear
till the leaves are shooting. The ♀-catkin in this instance is also
long and cylindrical. The cupule in _C. betulus_ is 3-lobed, and
to a slight extent only embraces the base of the ribbed nut (Fig.
331); each lobe corresponds to a floral-leaf. ~Whilst the carpels
are placed medianly in _Corylus_, in _Carpinus_, on the other hand,
they are situated transversely, as in the case of the _Betulaceæ_.
The lamina of the leaf is not conduplicate in the bud, but flat,
and folded only along the lateral veins, which are also indicated
in the form of the fully-developed leaf; otherwise the vegetative
characters are essentially the same as in the Hazel. The cotyledons
are aerial.--_Ostrya_ resembles the Hornbeam, but the cupule completely
envelopes the nut, as a sac open at the apex (Eur., N. Am., Japan).~
N. Am., Asia, and Europe; 25 species.--Fossil forms in the
Oligocene. Wind-pollinated. USES. As timber (_Carpinus betulus_)
and firewood. The fruits of _C. avellana_ (ordinary Hazel-nut),
_C. tubulosa_ (Lambert’s nut) and _C. colurna_ (Turkish Filbert)
are edible.
[Illustration: FIG. 330.--Diagrams of the ♂-flower (_A_) of
_Corylus_ and the dichasium of the ♀-flowers (_B_).]
[Illustration: FIG. 331.--Nut of the Hornbeam with cupule.]
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