=Mode of Life.= The Fungi have no chlorophyll, and are thus unable in
any stage of their existence to assimilate carbon; they must therefore
live as _saprophytes_ or _parasites_. There is, however, no strong line
of demarcation between these; many Fungi commence as true parasites,
but only attain their full development upon or in dead plants or
animals (_Rhytisma_, _Empusa_). Many saprophytes may occasionally
appear as parasites, and are then designated “_facultative parasites_”
(_Nectria cinnabarina_, _Lophodermium pinastri_), in contradistinction
to those which only appear as parasites, “_obligate parasites_”
(Mildew, Brand-and Rust-Fungi, _Cordyceps_).
The parasites which live on the surface of the host-plant are termed
_epiphytic_ (Mildew, _Fusicladium_); and those living in its tissues
are termed _endophytic_ (_Ustilago_, _Peronospora_). _Epizoic_
(_Oidium tonsurans_, _Laboulbenia_) and _endozoic_ Fungi (_Cordyceps_,
_Entomophthora_), are distinguished, in the same manner, as those
which live on the surface or in the interior of animals. The Fungi
designated _pathogenic_ are especially those which produce disease in
human beings and in animals.
Most of the diseases of plants are attributed to the parasitic Fungi.
These force their way into the host-plant by piercing the outer wall
of the epidermis, as in the Potato-disease; or by growing in through
the stomata, _e.g._ the summer generations of the Rust of Wheat; or
they can only penetrate through a wound, _e.g. Nectria_. Some effect
an entrance into the host-plant by the secretion of a poisonous matter
or ferment, which softens and destroys the cell-walls (_Sclerotinia_).
Some Yeast and Mould Fungi secrete ferments (enzymes), which, for
example, convert cane-sugar into a sugar capable of fermentation.
The relation of the parasitic Fungus to the host-plant is mainly of two
kinds. In the one case, the cell-contents are destroyed, the protoplasm
is killed, and the cellular tissue becomes discoloured and dies
(_Peronospora_, _Armillaria mellea_, _Polyporus_); in the other case,
the parasite has an irritating effect on the cellular tissue, whereby
the affected organ grows more rapidly and becomes larger than normal,
producing _hypertrophy_. Such malformations are termed _Fungi-galls_
(Mycocecidia); in this manner “witches’ brooms” are produced by
_Æcidium_, “pocket-plum” by _Taphrina_, and other deformities by
_Exobasidium_ and _Cystopus candidus_. This hypertrophy may either be
produced by a vigorous cell-multiplication, which is most frequently
the case, or by the enlargement of the individual cells (_Synchytrium_,
_Calyptospora_). The relation between host and Fungus among the Lichens
is of a very peculiar nature, termed “_symbiosis_.”
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