In the genera _Thamnidium_ and _Chætocladium_ the gradual
diminution of the sporangia, and the reduction of the number
of spores can be distinctly followed. In _Thamnidium_ the
number of spores is often reduced to one, which is _free_ in
the sporangium. In _Chætocladium_ however the sporangia are
typically _one-spored_, the spore is always united with the
sporangium, and the two become a single body, the so-called
_conidium_, which is in reality a closed sporangium. How
close is the connection between the sporangia and conidia of
_Thamnidium_ and _Chætocladium_, is seen from the fact that,
in the conidial stage of _Chætocladium_ the same whorl-form of
branching appears as in the sporangial stages of _Thamnidium
chætocladioides_, and also, that the conidia of _Ch.
fresenianum_ throw off the former sporangium-wall (exosporium),
while _Ch. jonesii_ germinates without shedding its exosporium.
The Phycomycetes have doubtless sprung from Water-Algæ and
inherit the sporangia from them. On this supposition, as the
Phycomycetes assumed a terrestrial mode of life, the sporangia
would become adapted to the distribution of the spores by
means of the air, the sporangia would become small, contain
dust-like spores, and would eventually become closed-sporangia,
_i.e._ conidia. The conidia are a terrestrial method for the
multiplication of Fungi. In the Hemiasci and the Ascomycetes the
sporangia are still preserved, but in every instance they are
adapted to terrestrial spore-distribution, their spores being
set free on the destruction of the sporangium-wall (generally
shot out) and distributed through the air. For further examples
of spore-distribution see below, p. 91–93.
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